Cochlis transgrediens ( Schaffer, 1912 )

Cochlis transgrediens ( Schaffer, 1912)

Figs 17A–D View FIGURE 17

* Natica transgrediens Schaff.— Schaffer 1912: 166 , pl. 54, figs 8–11.

N [ atica]. ( N [ acca].) transgrediens Schff.— Sieber 1958: 140 .

Natica millepunctata transgrediens Schaffer 1912 — Hölzl 1958: 212, pl. 19, figs 5–5a.

Natica millepunctata Lamarck— Răileanu & Negulescu 1964: 175 View in CoL , pl. 13, fig. 11 [ non Natica millepunctata Lamarck, 1822 View in CoL = Naticarius stercusmuscarum ( Gmelin, 1791) View in CoL ].

Natica ( Nacca) transgrediens Schaffer, 1912 — Steininger et al. 1971: 395, pl. 7, figs 11–12.

Type material. Lectotype (designated herein): GBA 1912 /004/001/3, SL: 23.1 mm, MD: 23.5 mm, Mörtersdorf ( Austria), illustrated in Schaffer (1912: pl. 54, fig. 10) . Paralectotypes: GBA 1912 /004/0017/1 SL: 24.7 mm, MD: 24.5 mm, Maria Dreieichen ( Austria), illustrated in Schaffer (1912: pl. 54, fig. 8) . GBA 1912 /004/0017/2, Maria Dreieichen ( Austria), illustrated in Schaffer (1912: pl. 54, fig. 9).

Illustrated material. NHMW 1934/0001/0123a, SL: 22.5 mm, MD: 22.9 mm, Loibersdorf ( Austria), Figs 17B View FIGURE 17 1 –B View FIGURE 1 4 View FIGURE 4 . NHMW 1934/0001/0123b, SL: 25.2 mm, MD: 23.5 mm, Loibersdorf ( Austria), Figs 17A View FIGURE 17 1 –A View FIGURE 1 4 View FIGURE 4 . Coll. Anton Breitenberger, SL: 34.2 mm, MD: 31.9 mm, Mörtersdorf ( Austria), Figs 17C View FIGURE 17 1 –C View FIGURE 1 4 View FIGURE 4 . Coll. Anton Breitenberger, SL: 28.2 mm, MD: 31.5 mm, Mörtersdorf ( Austria), Figs 17D View FIGURE 17 1 –D View FIGURE 1 4 View FIGURE 4 .

Additional material. 5 spec., NHMW 1848 View Materials /0003/0101, Loibersdorf ( Austria) ; 6 spec., NHMW 1849 View Materials /0013/0019, Mörtersdorf ( Austria) ; 7 spec., NHMW 1849 View Materials /0013/0020, Molt ( Austria) ; 1 spec., NHMW 1870 View Materials /0054/0031, Molt ( Austria) ; 1 spec., NHMW 1860 View Materials /0050/0309, Eggenburg ( Austria) .

Revised description. Medium sized, robust, globose shell of ~3.5 teleoconch whorls ( SL /MD = ~1.0). Protoconch unknown. Spire low conical, slightly gradate; apical angle 120–130°. Suture linear, distinctly incised. Teleoconch whorls with moderately wide, weakly convex subsutural shelf, delimited by weak shoulder angulation. Last whorl moderately expanding, attaining ~91–95% of total height. Periphery regularly convex, placed mid-whorl (~51%). Growth lines moderately distinct and weakly prosocyrt in apical view, weak at periphery. Aperture widely D-shaped, prosocline. Position of adapical tip of aperture high (~85%), distinctly below shoulder, above mid-whorl. Aperture attaining ~75% of total height. Columellar lip broad, strongly reinforced, basal lip weakly expanded, thickened, outer lip thin. Columellar angle ~20–25°. Marked opercular ridge in outer lip. Umbilicus semicircular, wide (~17%), deep with prominent growth lines. Parietal callus moderately long (~46%), weakly thickened, with concave margin, not expanding over base. Anterior lobe of parietal callus indistinct, partly fused with funicle. Funicle very broad, depressed, not sharply delimited, forming relatively narrow, shield-like umbilical callus. Adapical and abapical sulci shallowly concave. Basal fasciole rounded with prominent growth lines. No color pattern preserved. Operculum unknown.

Discussion. This is a very common Naticidae in the lower Eggenburgian Loibersdorf Formation in the Eggenburg region in Austria. We did not find any closely related coeval species from the Circum-Mediterranean region. Cochlis degrangei ( Cossmann & Peyrot, 1919) , from the Early Miocene of France, differs in the elongate shell and broad funiculus, which fills a large part of the umbilicus (see Robba et al. 2016: pl. 1, figs 3–5). The Early Miocene Cochlis neglecta ( Mayer, 1858) differs in its prominent, narrow, but well delimited funicle (see Robba et al. 2016: pl. 2, figs 9–15). Similarly, the Early Miocene Cochlis tigrina ( Defrance, 1825) differs in its prominent funicle and broad, deep umbilical channel (see Robba et al. 2016: pl. 4, figs 1–5). The Badenian Cochlis cuniculata nov. sp. is slightly reminiscent in shape and umbilical features but has a much narrower columella lip, the parietal callus is hardly developed, an even weaker funicle and a less expanding last whorl.

Paleoenvironment. The Loibersdorf Formation formed in shallow sublittoral environments with sand bottoms in few meters water depth ( Mandic et al. 2004).

Central Paratethys. Eggenburgian (Early Miocene): North Alpine Foreland Basin: Kaltenbachgraben ( Germany) ( Hölzl 1958); Gauderndorf, Loibersdorf, Mörtersdorf, Maria Dreieichen, Eggenburg ( Schaffer 1912). Transylvanian Basin: Coruş ( Romania) ( Răileanu & Negulescu 1964).