Cochlis neglecta ( Mayer, 1858 )

Cochlis neglecta ( Mayer, 1858) View in CoL

Figs 12A–D View FIGURE 12

* Natica neglecta Mayer— Mayer 1858: 388 View in CoL , pl. 11, fig. 2.

Natica epiglottina Lam. var. moldensis Schaff.— Schaffer 1912: 165 , pl. 54, figs 1–4.

Natica neglecta Mayer-Eymar View in CoL — Cossmann & Peyrot 1919: 403, pl. 11, figs 11–14.

Natica ( Natica) neglecta Mayer, 1858 View in CoL — Glibert 1952a: 76, pl. 6, fig. 4.

Natica ( Natica) neglecta Mayer, 1858 View in CoL — Glibert 1952b: 259, pl. 2, fig. 4.

N [ atica]. ( N [ acca].) epiglottina View in CoL moldensis Schff.— Sieber 1958: 140.

Polinices ( Naticina) epiglottina moldensis ( Schaffer 1912) — Hölzl 1958: 206, pl. 19, fig. 7.

Natica neglecta Mayer 1858 View in CoL — Janssen 1969: 173, pl. 5, figs 18–24, text-fig. 12.

Natica ( Nacca) epiglottina moldensis Schaffer, 1912 — Steininger et al. 1971: 395, pl. 7, fig. 10. Naticarius neglectus Mayer— Nordsieck, 1972: 71 , pl. 17, fig. 90.

Natica neglecta Mayer, 1858 View in CoL — Janssen 1984: 202, pl. 57, figs 5–7, 12.

Natica neglecta Mayer, 1859 View in CoL — Lozouet et al. 2001: 43, pl. 18, fig. 2.

Natica neglecta Mayer, 1858 View in CoL — Wienrich 2001: 432, pl. 70, fig. 1, pl. 87, figs 3–4, pl. 88, fig. 7.

Cochlis neglecta ( Mayer, 1858) View in CoL — Moths et al. 2010: 37, pl. 11, fig. 7.

Cochlis neglecta ( Mayer, 1858) View in CoL — Robba et al. 2016: pl. 2, figs 9–15.

Cochlis neglecta (Mayer, 1859) View in CoL — Stein et al. 2016: 46, text-fig. 18, pl. 22, fig. 2.

? Cochlis sp. — Kovács et al. 2023: fig 30/38.

non Natica epiglottina moldensis Schaffer, 1812 [ sic]– Steininger 1973: 412, pl. 4, fig. 12 [= Cochlis sp. ].

Type material. Syntype, ETH Zürich ( Switzerland), SL: 23 mm, MD: 21 mm, ‘environs de Bordeaux’ ( France), Aquitanian, Early Miocene, illustrated in Mayer (1859: pl. 11, fig. 2) ,

Illustrated material. Lectotype of Natica epiglottina moldensis Schaffer, 1912 (designated herein): GBA 1912/004/15/1, SL: 23.0 mm, MD: 23.2 mm, Maria Dreieichen ( Austria), illustrated in Schaffer (1912: pl. 54, fig. 1), Figs 12A View FIGURE 12 1 –A View FIGURE 1 4 View FIGURE 4 . Paralectotypes: GBA 1912/004/15/1, SL: 18.3 mm, MD: 16.6 mm, Maria Dreieichen ( Austria), illustrated in Schaffer (1912: pl. 54, fig. 2), Figs 12B View FIGURE 12 1 –B View FIGURE 1 4 View FIGURE 4 . GBA 1912/004/15/1, SL: 15.6 mm, MD: 14.4 mm, Maria Dreieichen ( Austria), illustrated in Schaffer (1912: pl. 54, fig. 3), Figs 12C View FIGURE 12 1 –C View FIGURE 1 4 View FIGURE 4 . GBA 1912/004/15/1, SL: 17.8 mm, MD: 15.8 mm, Maria Dreieichen ( Austria), illustrated in Schaffer (1912: pl. 54, fig. 4), Figs 12D View FIGURE 12 1 –D View FIGURE 1 4 View FIGURE 4 .

Revised description. Medium sized, solid, elongate-ovate shell; higher than wide ( SL /MD = ~1.1). Spire high conical, of convex whorls; apical angle 95–100°. Protoconch poorly preserved. Suture linear, adpressed. Last teleoconch whorl high, ovate, with steep, indistinct subsutural ramp, delimited by indistinct, rounded shoulder angulation. Last whorl moderately expanding, not inflated, attaining ~90% of total height. Periphery placed below mid-whorl (~45%), regularly convex. Growth lines distinct and prosocyrt in apical view, delicate and prosocyrt at periphery. Aperture moderately wide, D-shaped, with maximum diameter in abapical third. Position of adapical tip of aperture moderately low (~76%), far below shoulder. Aperture attaining ~78–80% of total height. Columellar lip weakly reinforced, basal lip and outer lip thin. Columellar angle ~26–32°. Opercular ridge in outer lip forming broad, low swelling. Umbilicus moderately wide (~16%), deep, partly filled by funicle. Parietal callus long (~51%), thickened, not expanding over base with straight to weakly concave margin. Small, semicircular anterior lobe. Funicle prominent, cord- to pillar-like, narrow, separated from umbilical wall by broad umbilical channel. Small but prominent semicircular umbilical callus slightly at or slightly below middle of columellar lip well separated from anterior lobe. Basal fasciole rounded to weakly angled at aperture. Color pattern not preserved. Operculum unknown from Paratethyan occurrences (see Robba et al. 2016: 131 for description of operculum and protoconch).

Paratethyan synonyms. Natica epiglottina moldensis Schaffer, 1912 cannot be distinguished from Cochlis neglecta and therefore, is treated as subjective junior synonym of Natica neglecta Mayer, 1858 . Most probably, Schaffer (1912) was not aware of the paper by Mayer (1858).

Discussion. Cochlis degrangei ( Cossmann & Peyrot, 1919) , from the Early Miocene of France and Italy, is comparable in outline and umbilical features, but differs from Cochlis neglecta ( Mayer, 1858) in its much larger umbilical callus (see Robba et al. 2016: pl. 1, figs 3–4). The Eocene Cochlis epiglottina ( Lamarck, 1804) , with which this species was compared by Schaffer (1912), differs in its lower spire, larger umbilical callus and wider umbilical channel (see Robba et al. 2016: pl. 1, fig. 13a, pl. 2, figs 1–2). Cochlis infelix ( Sacco, 1890) , from the Rupelian of Sassello ( Italy), might be a closely related species, but differs in its gradate spire (see Robba et al. 2016: pl. 2, fig. 6). A specimen from the Karpatian of Várpalota ( Hungary), illustrated by Steininger (1973: pl. 4, fig. 12) as Natica epiglottina moldensis , differs in its larger umbilical callus and broader funiculus and represents another Cochlis species. A specimen from the Ottnangian of Hungary, described by Kovács et al. (2023) might represent Cochlis neglecta but the preservation does not allow a clear identification.

Paleoenvironment. Coastal marine, lagoonal in vicinity of mudflats and oyster reefs ( Mandic et al. 2004).

Distribution in Central Paratethys. Eggenburgian (Early Miocene): North Alpine Foreland Basin: Kaltenbachgraben ( Germany) ( Hölzl 1958); Eggenburg, Maria Dreieichen, Mörtersdorf, Mold, Nonndorf ( Austria) ( Steininger et al. 1971).? Ottnangian (Early Miocene): Early Miocene North Hungarian Basin: Parádfürdö ( Hungary) ( Kovács et al. 2023).

North Sea. North Sea Basin: Burdigalian/Langhian (Early/Middle Miocene): Winterswijk Miste ( the Netherlands) ( Janssen 1984) Dingden, Kevelaer, Werder ( Germany) ( Janssen 1969; Moths et al. 2016).

Northeastern Atlantic. Aquitanian–Burdigalian (Early Miocene): Aquitaine Basin: ( France): La Houga, Léognan, Lucbardez-et-Bargues, Mérignac, Saint-Martin-d’Oney, Saucats, Villandraut ( France) ( Cossmann & Peyrot 1919; Robba et al. 2016).

Proto-Mediterranean Sea. Burdigalian (Early Miocene): Colli Torinesi : Calle Ceppi, Val Sanfrà ( Italy); Langhian (Middle Miocene): Colli Torinesi : Monte dei Cappuccini ( Italy); Tortonian (Late Miocene): Po Basin: Borelli, Montegibbio, Rio di Bocca d’Asino, Sant’Agata Fossili ( Italy) ( Robba et al. 2016).