Naticarius lucapedrialii, Harzhauser & Landau & Guzhov, 2025

Naticarius lucapedrialii nov. sp.

Figs 23A–B View FIGURE 23

? Natica ( Cochlis) dillwyni Payr.— Boettger 1902: 81 View in CoL [ non Notocochlis dillwynii ( Payraudeau, 1826) View in CoL ].? Natica ( Cochlis) dillwyni Payr. var. plicatula Bronn.— Boettger 1906: 89 View in CoL [ non Notocochlis dillwynii ( Payraudeau, 1826) View in CoL ]. Natica ( Euspira) catena Da View in CoL Costa var. helicina Brocchi, 1814 View in CoL — Moisescu 1955: 208, pl. 19, figs 14–15 [ non Euspira helicina

( Brocchi, 1814)]. Polinices ( Euspira) catena var. helicina ( Brocchi 1814) — Korobkov 1955: pl. 41, figs 23–24 [ non Euspira helicina ( Brocchi,

1814)]. Natica ( Nacca) dillwyni var. plicatula (Bronn 1831) —Kojumdgieva in Kojumdgieva & Strachimirov 1960: 119, pl. 33, fig. 7. Natica sp. —Caze et al. 2014: 33, figs 5A1–A3.

Type material. Holotype, NHMW 2024 View Materials /0240/0001, SL: 16.9 mm, MD: 18.2 mm, Lăpugiu de Sus ( Romania), Figs 23A View FIGURE 23 1 –A View FIGURE 1 6 View FIGURE 6 . Paratype: MNHN A31069 View Materials , SL: 16.5 mm, MD: 16.0 mm, Lăpugiu de Sus ( Romania), illustrated in Caze et al. (2014: figs 5A1–A3), photo by Philippe Loubry ( MNHN /CNRS), Figs 23B View FIGURE 23 1 –B View FIGURE 1 6 View FIGURE 6 .

Type locality. Lăpugiu de Sus ( Romania), Făget Basin .

Type stratum. Silt and clay of the Dej Formation.

Age. Middle Miocene, Badenian (Langhian).

Etymology. In honor of Luca Pedriali (San Martino, Ferrara, Italy) in recognition of his contributions to the study of Cainozoic Naticidae faunas. We note that this name should not be confused with Cochlis pedrialii Van Dingenen, Ceulemans & Landau, 2016 from the Lower Pliocene of northwestern France, also dedicated to this outstanding naticid expert.

Diagnosis. Small, depressed globose shell, low spire, with moderately wide umbilicus and prominent, cord-like, narrow, strongly raised funicle; color pattern of small, vaguely diamond-shaped, close-set scales arranged in three bands.

Description. Small, depressed globose shell of ~3.5 teleoconch whorls; only slightly wider than high (SL/ MD = ~0.9). Spire low conical with moderately convex whorls; apical angle ~130°. Protoconch unknown. Suture linear, adpressed. Teleoconch whorls with moderately broad, weakly concave subsutural shelf, delimited by angulate shoulder. Last whorl regularly convex below shoulder, moderately expanding, attaining ~95% of total height. Periphery placed mid-whorl (~49%). Growth lines delicate, weakly prosocyrt in apical view, subobsolete at periphery.Aperture moderately wide, ovate D-shaped. Position of adapical tip of aperture high (~86%), not far below shoulder, distinctly above mid-whorl. Aperture attaining ~82% of total height. Columellar lip weakly reinforced, basal lip slightly thickened, outer lip thin. Columellar angle 15–23°. Opercular ridge in outer lip insignificant. Umbilicus moderately wide (~17%), deep, with delicate growth lines. Parietal callus very long (~61%), weak, not expanded over base. Anterior lobe absent (or broken off in both available specimens). Funicle prominent, cord-like, narrow, strongly raised, terminating in abapical third of columellar lip in small, prominent, subtrigonal umbilical callus. Broad, concave adapical sulcus, narrower, deeply concave abapical sulcus delimited by weak spiral ridge. Basal fasciole weak, rounded. Color pattern of small, vaguely diamond-shaped, close-set dark scales arranged in three broad bands separated by two narrow, light bands placed slightly above periphery and at base with fewer scales. Operculum unknown.

Discussion. According to Robba et al. (2016: 121), species of the genus Naticarius have a teleoconch similar to those of the genus Cochlis , but their opercula have the outer surface with many ribs instead of two or three. Therefore, without the operculum we cannot ascribe species to one or other genus with certainty. However, we place this species in Naticarius based on its narrow and strongly raised funicle, which is comparable to the extant Mediterranean Naticarius hebraeus (Martyn, 1786) and Naticarius stercusmuscarum ( Gmelin, 1791) . Naticarius lucapedrialii nov. sp. displays a very intense color pattern under UV-light, which differs from the Paratethyan Cochlis species in its small, diamond-shaped scales and spiral bands. This pattern is reminiscent of that of N. hebraeus , which however, does not form bands. Cochlis raropunctata (Sasso, 1827) , from the Tortonian of Italy, has a similar funicle but is ovate-globose and differs in its color pattern of circular dots (see Pedriali & Robba 2005: pl. 4, figs 1–6, pl. 9, figs 14–19; Chirli 2008: pl. 23, figs 13–18, pl. 24, figs 4–13; Robba et al. 2016: pl. 3, fig. 4). The umbilical features of Naticarius lucapedrialii nov. sp. are superficially reminiscent of those of Cochlis pseudovittata Robba, Pedriali & Quaggiotto, 2016 , from the Tortonian of Italy, but the Italian species differs in its larger umbilical callus and color pattern of regularly spaced dots (see Robba et al. 2016: pl. 2, fig. 16). Kojumdgieva in Kojumdgieva & Strachimirov (1960) confused this species with the extant Mediterranean Notocochlis dillwynii ( Payraudeau, 1826) ], which differs in its longer, elongate semicircular umbilical callus and trigonal anterior lobe.

Paleoenvironment. Unknown.

Distribution in Central Paratethys. Badenian (Middle Miocene): Făget Basin: Coşteiu de Sus, Buituri, Lăpugiu de Sus ( Romania) ( hoc opus; Boettger 1906; Moisescu 1955); Dacian Basin: Bivolare ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960).

Genus Tectonatica Sacco, 1890

Type species. Natica tectula Sacco, 1890 View in CoL ; by monotypy in Sacco (1890: 33). Pliocene, Italy.