Pseudolinices pseudoredemptus ( Friedberg, 1923 ), 2025

Pseudolinices pseudoredemptus ( Friedberg, 1923) nov. comb.

Figs 30A–D View FIGURE 30

* Natica View in CoL ( Poliniceps [ sic]) pseudoredempta Friedb.— Friedberg 1923: 434 , pl. 26, fig. 12, text-fig. 78.

Polynices redempta Micht.— Strausz 1954: 23 , pl. 6, figs 131a–d [ non Pseudolinices redempta ( Michelotti, 1847) ].

Polinices ( Polinices) pseudoredempta ( Friedberg 1923) —Kojumdgieva in Kojumdgieva & Strachimirov 1960: 120, pl. 33, figs 8–9.

? Polynices ( Lunatia) ex gr. proredempta Sacco— Kókay 1966: 55 , pl. 7, fig. 7.

? Polynices redempta dertoconvexa Sacco— Kókay 1966: 55 , pl. 7, fig. 8 [ non Pseudolinices redemptus ( Michelotti, 1847) ]. Polynices pseudoredempta (Friedberg) — Kókay 1966: 55, pl. 7, fig. 9.

Natica ( Polynices) redempta Michelotti, 1847 View in CoL — Strausz 1966 ( pars): 231, pl. 47, figs 19–31 [ non Pseudolinices redempta ( Michelotti, 1847) ].

Polinices ( Polinices) pseudoredemptus ( Friedberg, 1923) —Hinculov in Iliescu et al. 1968: 135, pl. 32, fig. 17.

Polinices ( Polinices) redempta Micht.— Popović & Gagić 1969: 99 View in CoL , pl. 8, fig. 12.

Polinices pseudoredemptus ( Friedberg, 1923) — Bałuk 1995: 197, pl. 15, fig. 6.

Euspira catena helicina (Brocchi) View in CoL — Popa & Chira 1999: 400, pl. 3, fig. 8 [ non Euspira helicina ( Brocchi, 1814) View in CoL ].

Polinices pseudoredemptus ( Friedberg, 1923) — Harzhauser 2002: 87, pl. 5, figs 1–4.

Polinices pseudoredemptus ( Friedberg, 1923) — Bałuk 2006: 206, pl. 3, fig. 7.

non Polinices pseudoredempta (Friedberg) — Ionesi & Nicorici 1994: 58, pl. 2, fig. 11 [= Euspira sp. ].

non Polinices cf. pseudoredemptus ( Friedberg, 1923) — Höltke 2009: 88, fig. 34 [= Euspira sp. ].

Type material. Lectotype (designated herein), SL: 18.1 mm, MD: 13.5 mm, Korytnica ( Poland), illustrated in Friedberg (1923: pl. 26, fig. 12).

Illustrated material. NHMW 2024/0247/0001, SL: 12.2 mm, MD: 10.7 mm, Kleinebersdorf ( Austria), Figs 30A View FIGURE 30 1 –A View FIGURE 1 4 View FIGURE 4 . NHMW 2024/0247/0003, SL: 12.8 mm, MD: 11.0 mm, Kleinebersdorf ( Austria), Figs 30B View FIGURE 30 1 –B View FIGURE 1 4 View FIGURE 4 . NHMW 2024/0247/0004, SL: 12.3 mm, MD: 10.2 mm, Kleinebersdorf ( Austria), Figs 30C View FIGURE 30 1 –C View FIGURE 1 4 View FIGURE 4 . NHMW 2024/0247/0005, SL: 10.8 mm, MD: 8.9 mm, Kleinebersdorf ( Austria), Figs 30D View FIGURE 30 1 –D View FIGURE 1 4 View FIGURE 4 .

Additional material. 55 spec., NHMW 1861 View Materials /0050/0156, Kleinebersdorf ( Austria) ; 10 spec., NHMW 1846 View Materials /0037/0440, Kleinebersdorf ( Austria) ; 1 spec., NHMW 1859 View Materials /0045/0049, Grossrussbach ( Austria) ; 2 spec., NHMW 1861 View Materials /0050/0036, Rückersdorf ( Austria) ; 4 spec., NHMW 1859 View Materials /0045/0262, Weinsteig ( Austria) ; 70 spec., NHMW 2024 View Materials /0248/0001, Karnabrunn ( Austria) .

Revised description. Small, solid, elongate ovate shell with high spire; higher than wide ( SL /MD = 1.1– 1.3). Spire high turbiniform, with strongly convex whorls; apical angle 95–140°. Protoconch low turbiniform, of 2.5 smooth, convex whorls. Suture linear, adpressed. Teleoconch whorls with narrow, convex subsutural ramp, delimited by rounded shoulder. Last whorl allometrically expanding, basally inflated, attaining ~95% of total height. Periphery far below mid-whorl (~38%). Growth lines delicate, weakly prosocyrt in apical view, delicate, prosocline at periphery. Aperture wide, D-shaped. Position of adapical tip of aperture low (~78%), far below shoulder, below mid-whorl. Aperture attaining ~75% of total height. Columellar lip slightly strongly reinforced, basal lip slightly thickened, outer lip thin. Columellar angle ~35°. Opercular ridge in outer lip not developed. Umbilicus fully sealed by very prominent, thickened, pad-like, often strongly convex callus. Parietal callus very long (~80%), strongly thickened with prominent adapical swelling, slightly expanding, with straight margin. Anterior lobe fused with parietal and umbilical callus. Basal fasciole solid, rounded to weakly angled. Color pattern and operculum unknown.

Discussion. Pseudolinices pseudoredemptus ( Friedberg, 1923) is a unique species due to its often strongly allometrically grown last whorl with the suture shifting far below the shoulder of the penultimate whorl. Its small size and elongate outline distinguish it distinctly from Pseudolinices empina ( De Gregorio, 1885) . In addition, its thick, pad-like umbilical callus is highly characteristic and separates it from other Circum-Mediterranean Neogene Pseudolinices species. Pseudolinices pseudoredemptus is quite variable in shape, which might be a result of its adaptation to strong environmental fluctuations (see below).

Paleoenvironment. Shallow marine, inner neritic. Occurrences from the Korneuburg Basin in Austria indicate mudflats of coastal lagoons ( Harzhauser 2002) and Bałuk (1995, p. 198) suggested that Pseudolinices pseudoredemptus was also a brackish-water species.

Distribution in Central Paratethys. Karpatian (Early Miocene): Korneuburg Basin: Helfens, Hipples, Karnabrunn, Kleinebersdorf, Leobendorf, Niederkreuzstetten, Rückersdorf, Stetten, Weinsteig, Wetzleinsdorf ( Austria) ( Harzhauser 2002; hoc opus). Badenian (Middle Miocene): Korytnica Basin: Korytnica, Małoszów ( Poland) ( Friedberg 1923; Bałuk 2006); Pannonian Basin: Herend-Marko, Várpalota ( Hungary) (Hörnes 1856; Strausz 1966); Timok Basin: Pivinčka Reka ( Serbia) ( Popović & Gagić 1969): Borod Basin: Borehole 575 at Cetea ( Romania) ( Popa & Chira 1999); Mehadia Basin : Valea Calvei, Valea Bela Reca ( Romania) (Hinculov in Iliescu et al. 1968); Dacian Basin: Dobrusha ( Bulgaria) (Kojumdgieva in Kojumdgieva & Strachimirov 1960).