Hystatoderes weissi Lameere, 1915

Hystatoderes weissi Lameere, 1915 View in CoL

( Fig. 1, 4 A-C, 6C, F & H)

Emphiesmenus weissi Lameere, 1915: 56 View in CoL (description).

Emphiesmenus weissi Lameere, 1919: 115 View in CoL (catalogued).

Emphiesmenus weissi Gressitt & Rondon, 1970: 18 View in CoL , Fig. 4, d-e (male description) [citation from Laos].

Emphiesmenus weissei Hua, 2002: 206 View in CoL (catalogue, misspelling) [citation from China].

Hystatoderes weissi Li et al., 2014: 83 View in CoL (catalogue, distribution in China).

Records from China and Laos will be treated in the second part of this revision.

Material examined

Type material studied

Holotype ♀, 41 mm ( Fig. 4 B-C, 6F &H): Museum Paris, Tonkin Sept., Frontière de Chine, Ha Giang, A. Weiss 1901 / oct.-dec. / Emphiesmenus WeissiA.Lameere vid. 1915/ TYPE( MNHN) / Holotype Emphiesmenus weissi Lameere, 1915 , vid. Drumont et al., 2023.

FAUNITAXYS - ISSN (Print): 2269 - 6016 - ISSN (Online): 2970 - 4960

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C, F, H. Hystatoderes weissi ( Lameere, 1915) . C. ♂ ( ADC), forebody. F . ♀ ( MNHN), forebody. H . ♀, basal part of elytra.

Other material studied

1 ♂, Vietnam, Ha Giang province, env. Lung Cu , 1200 m, local collector leg. ( ADC) ( Fig. 4A) .

Description

Male ( Fig. 4A, 6C)

Dimensions. – Body length: 31mm.

Habitus . – With a stocky appearance (like H. vitalisi ), oblong, concolor dark brown, less than twice as long as wide at the basal third of elytra. Pronotum, elytra, scutellum, head and antennae concolor, pronotum with a golden dense pilosity near apical and basal margins. Legs brown with elongated tarsi. Antennae barely exceeding the elytral half.

Head. – Robust, slightly wider than long; frons and vertex with a scattered punctation evenly distributed over the surface; median longitudinal groove on the top running between the antennal tubercles towards the rear and bordered by a strong carina; a small other carina bordering the upper part of ocular lobe joining the first one near the middle of the vertex. Eyes moderately indented; lower lobe barely wider than the upper lobe; inter-ocular space equal to the width of the upper lobe. Mandibles regularly rounded and strongly punctate, wider from base to central part, bifid with an intermediate tooth in the basal third.

Antennae. – Rather short, just reaching the apical third of the elytra. Scape strongly vermiculated and wrinkled, slightly flattened on dorsal side, enlarged at apex with rounded outer corner, antennomere 3 as long as scape, with irregularly spaced strong punctation; antennomere 4 punctate or with few longitudinal groove; antennomeres 5–10 equal in length and expanding bell-shaped towards the apex, with longitudinal grooves; last antennomere longer with bevelled apex.

Pronotum. – Transverse, hexagonal, swollen on the disc; surface strongly vermiculate with huge wrinkles, partially hairless except along the apical and basal margin, which have a fringe of long golden hairs; apical margin much wider than head; apical angles irregularly shaped with a fairly strong spine offset downward; a weaker tooth about halfway up the pronotum followed by a strong, slightly downwardfacing spine. Under this last tooth, the pronotum narrows to the basal border without teeth. Scutellum: large, rounded, granular and covered with sparse golden setae.

Elytra. – Almost smooth and hairless, showing only areas with weak fine wrinkles and a small punctuation on the basal part around the scutellum, punctuation becoming stronger on the humeral angles; costae inconspicuous; lateral margins relatively wide; apical angles weakly sharp, without spine.

Legs. – Dark brown; femora straight, punctate and covered with a scattered pilosity; tibiae punctate, strongly concave inner face and finely crenellated on the lower side, widening towards the apex and with few scattered hairs; apex toothed at both sides; tarsi narrow and elongated; onychium as long as tarsomeres 1- 3 combined.

Female ( Fig. 4 B-C, 6F & H)

Dimensions. – Body length holotype ♀: 41 mm.

Habitus . – Body moderately large, stout, thicken dorsal ventral side, and brighten red.

Head. – Remarkable shorter than wide. Eyes narrow in dorsal view, distantly separated, distance between eyes about haft of wide of the head and wider than dorsal lobe of the eyes. Labrum with some short setae. Mandible strong, robust, curved inward and downward, right mandible with inner apical teeth forming a bifid apex. Underside of head concave and not smooth.

Antennae. – Short, about 0.6 as long as body, 11-segmented; scape the thickest, as long as antennomere 3, round at sides, somehow little pressed in dorsal-ventral direction, expanded apically; antennomere 2 visible; antennomere 3 about as long as scape, mostly rounded but somehow flattened at ventral side, narrow at base and expanded apically; antennomeres 4 - 5 similar in shape and size, antennomeres 4 and 5 combined about as long as antennomere 3; antennomeres 6- 11 combined about as long as 3 -5 combined; antennomeres 1 -9 somehow smooth with punctuation; antennomeres 10 - 11 with wrinkles in the length.

Pronotum. – Short, about 0.6 as long as wide (measured between 2 tips of basal spines); apical margin concave; basal margin convex, making a basal lobe; surface smooth on the middle, disc convex and two lateral parts much more punctate, reaching the lateral margins; lateral margin with three evident spines and one fold forming a small spine, the basal one longest and more pointed that others. Except the smallest tooth, all spines are curved upward. Scutellum: short and rounded, making a semicircle with basal margin of pronotum, not smooth as the middle part of pronotum but less punctate than the elytra.

Elytra. – Reddish and shining, strong punctuation at base and reduced from the apical fourth; lateral margin curves up strongly from base and nearly to flatten at apex; apex rounded with a short angle.

Legs. – Flattened, same colour as the body, tarsi segment with lateral apical somehow point.

Diagnostic characters. – Hystatoderes weissi is easily distinguished from the other two species composing the genus by the following combination of characters:

- For the males (only one specimen studied): remarkable for the very transverse pronotum, marked with deep wrinkles and provided with three lateral spines, the vertex showing a strong longitudinal carina, and the blackish brown elytra, mat and with basal angles rounded and without tooth.

- For the females (only one specimen studied): by the relatively square pronotum with three evident spines and a fold of margin forming a small spine, and the reddish brown elytra with basal angles rounded and without tooth.

Etymology. – Described in honour of the collector A. Weiss.

Flight period / biotope altitude. – For H.weissi , only a few data are available, for the altitude the male was recorded at 1200 m and the flight period for the holotype female is October-December.

Distribution. – Extreme northern part of Vietnam ( Fig. 1)