Cyrto-Hypnum versicolor, (Hornsch. ex Mull. Hal.) W. R. Buck & H. A. Crum

Cyrto-hypnum versicolor (Hornsch. ex Müll. Hal.) W.R. Buck & H.A. Crum View in CoL Figs. 2–5 View Fig View Fig View Fig View Fig

Plants [small] or {minute}, green to light green, not glossy, forming a tiny patches on [rocks, soil and rotten wood] or {on rock surfaces}. Stem [ca. 15 mm long] or {ca. 0.5 mm long}, [loosely attached to the substrate] or {creeping on substrate, tightly attached by numerous rhizoids}, central strand absent, bipinnately branched; primary branches [to 4 mm long, moderately spaced, arranged on both sides of stem] or {2.5–3.0 mm long, densely arranged on one side of stem and arising vertically from it}; secondary branches [1.0– 1.2 mm long] or {0.5– 0.8 mm long}; surface of stem and branches [smooth] or {mammilose}, [stem covered with dense paraphyllia, on primary branches paraphyllia sparse or absent] or {paraphyllia sparse on stem and primary branches}, paraphyllia small, 40–45 µm long, narrow and truncate at apex or foliose. Stem leaves [regularly arranged] or {sparse due to dense branching pattern}, [ca. 0.5 mm long and 0.3 mm wide] or {ca. 0.25 mm long and 0.15 mm wide}, triangular, with narrow acumen filled by costa; costa strong, gradually narrowing distally, excurrent. Primary branch leaves [0.3–0.35 mm long and 0.15 mm wide] or {0.21– 0.27 mm long and 0.09–0.1 mm wide}, ovate-lanceolate, [acuminate, with obtuse apical cell] or {shortly acute or subobtuse at apex, with acute or obtuse apical cell}, shallowly keeled above; margins plane, densely serrate; costa single, gradually narrowing distally, ending well below apex, smooth, looking pale in contrast with dim lamina areolation; lamina cells uniform within the whole leaf, subquadrate and short rectangular, 5–8 µm long and 4–6 µm wide, moderately thick-walled, with single high papilla on dorsal leaf surface or, at places, on both surfaces. Secondary branch leaves ovate, 0.1–0.18 mm long, 0.05–0.06 mm wide, [acute] or [rounded at apex], with plane margins, serrate, lamina areolation like in primary branch leaves. Autoicous, sporophytes [not seen] or {numerous}.{Perichaetia large, whitish, perichaetial

leaves ca. 2.3 mm long, with oblong base and long triangular apex, not plicate, margins plane, finely serrulate at places, costa strong, narrowing distally, excurrent. Seta ca. 1 cm long, reddish-brown, smooth, straight or slightly bent shortly below the capsule. Capsules short, slightly curved, with ovate urn and short neck, urn ca. 0.9 mm long and 0.5 mm, operculum conic, with oblique beak ca. 0.55 mm long, annulus revoluble, consisting of two rows of inflated cells. Spores 7–9(–11) µm, smooth. Calyptra not seen}.

Specimens examined: Primorsky Territory, Slavyansky District, Suputinsky [Ussurijsky] Reserve, mixed conifer and broadleaved forest with hornbeam on north-faced slope, IX.1982, Vasiljeva & Ardeeva s.n. (LE). Sakhalinskaya Province, Kuril Islands, Kunashir Island, Mechnikova Mt. (43°58’27’’N – 145°42’48’’E), western foot of hill; forest with Picea glehnii , P. yezoensis and Abies sachalinensis , with dense undegrowth of Sasa sp. , mesic, moderately dark; on shaded side of big boulder, 4.VIII.2015, Koroteeva 15-3/2-7 (MHA).