Pantolyta melniki, Chemyreva & Kolyada, 2021

Pantolyta melniki sp.nov.

( Figs 2C, 4C, E, 6D, 8B, 9D, 10F, 11B, 13I, 14C)

Holotype. Female; Russia, Sakhalin Prov., Kunashir I., vicinity of Grozovoe , 8–15.VIII.2008, I. Melnik leg. ( ZIN).

Paratype. Republic of Korea, Gangwon Prov., Odaesan National Park, Dongsan-Li, near Woljeongsa , fir forest, Malaise trap, 19.VII–18.VIII.2003 [collector unknown], 1 male ( ZIN) .

Description. Female. Body length 3.5 mm; fore wing length 2.7 mm; antenna length 2.1 mm.

Coloration. Head, mesosoma and petiole dark brown; antenna, mandibles, venation and metasoma (except petiole) pale brown; legs and palpi yellowish brown.

V.G. Chemyreva & V.A. Kolyada. Taxonomy of Acropiesta , Anommatium , Erasikea and Pantolyta

Fig. 8. Pantolyta and Anommatium , female (A–E) and male (F). Head, dorsal view. A, P. dichromia sp. nov.; B, P. melniki sp. nov.; C, P. sciarivora ; D, P. radialis ; E, P. nigrocincta ; F, A. ashmeadi .

Head not nasiform, in dorsal view 1.25 times as wide as long (Fig. 8B), wider than mesosoma, smooth, with few scattered long setae. Temples just behind eyes parallel, broadly receding posteriorly. Occipital flange narrow, foveolate (Fig. 8B). Head in lateral view 1.21 times as high as long. Malar space 0.90 times as long as largest diameter of eye. Eye pubescent. Antennal shelf in frontal view with fine coriaceous sculpture below toruli. Face smooth, with long scattered setae. Head in frontal view without mouth conus; genae slightly convex and converging toward mouthparts. Clypeus transverse, 0.67 times as high as wide. Tentorial pits large and transverse. Labrum small, semicircular, hardly discernible ( Fig. 2C). Mandibles simple.

Antenna slender, weakly broadened apically ( Fig. 4C, E). Ratios of length to width of A1–A 15 in dorsal view: A1 (40: 8); A2 (12: 7); A3 (18: 7); A4 (12: 7); A5 (11: 7); A6 (11: 7); A7 (10: 7); A8 (10: 7); A9 (10: 7); A10 (10: 8); A11 (9: 9); A12 (9: 9); A13 (9: 9); A14 (9: 10); A15 (16: 11).

Mesosoma slightly compressed, 1.08 times as high as wide, in dorsal view 1.83 times as long as wide. Sides of pronotum bare, pronotal collar with scattered long setae; epomia strongly prominent; pronotal pit bare, with epomia inside (Fig. 9D). Mesoscutum slightly elongate, 1.13 times as long as wide, convex, with few scattered semierect setae. Notauli deep and complete, sculptured. Anterior scutellar pit deep and rounded. Scutellum large, widened posteriorly. Axillar depression scarcely pubescent, with two verriculate tubercles (Fig. 11B). Mesopleuron scarcely pubescent ventrally and anterodorsally, with subalar bridge posterodorsally and epicnemial bridge anteroventrally; epicnemial pit deep and pubescent; mesopleural pit developed as shallow depression (Fig. 10F).

V.G. Chemyreva & V.A. Kolyada. Taxonomy of Acropiesta , Anommatium , Erasikea and Pantolyta

Fig. 9. Pantolyta , female. Pronotum, lateral view. A, P. flexinervis ; B, P. sciarivora ; C, P. seticornis ; D, P. melniki sp. nov.; E, F, P. radialis .

Metascutellum narrow, pubescent; dorsellum distinct, with three short longitudinal keels. Propodeum transverse, 0.38 times as long as wide, entirely pubescent, with simple median keel; upper and lower posterior propodeal projections forming large teeth (Figs 11B and 13I). Sides of propodeum entirely pubescent, with three longitudinal keels (Fig. 10F). Legs slender.

Fore wing with closed radial cell (Fig. 14C); distance from marginal vein to basal vein / marginal vein length / radial cell length as 12 / 19 / 18.

Petiole of metasoma transverse 0.75 times as long as wide, with coarse rugose sculpture and few scattered setae laterally (Fig. 11B); petiole ventrally with several bunches of setae. T2 anteriorly with short striation, median groove the longest; T3–T6 narrow, scarcely pubescent; T7 and T8 long, strongly compressed laterally. S2 with short grooves at base and with numerous erect scattered setae; S3–S5 narrow, with scattered setae; S6 distinctly elongate and compressed, with few scattered setae.

Male. Body length 2.9 mm. Similar to female except following characters: antenna filiform; antennomeres cylindrical, with uniform short pubescence; A1 slender, 5.0 times as long as wide and 0.64 times as long as head width; A3 deeply emarginate, with keel extending to 0.6 of A3 length ( Fig. 6D); ratios of length to width of A1–A 5 in lateral view: A1 (40: 8); A2 (14: 8); A3 (29: 9); A4 (20: 7); A5 (18: 7); petiole elongate, 1.38 times as long as wide (Fig. 13I); T7 transverse, not compressed; S2–S7 with numerous semierect scattered setae; T8 and S8 subtriangular, rounded apically.

Fig. 10. Pantolyta , female (A–E) and male (F). Mesosoma and anterior part of metasoma, lateral view. A, P. micans ; B, P. nigrocincta ; C, P. rufiventris ; D, P. dichromia sp. nov.; E, P. simplicior sp. nov.; F, P. melniki sp. nov. Green arrow – upper posterior propodeal projection; white arrow – lower posterior propodeal projection; red arrows – median propodeal keel.

Comparison. Pantolyta melniki sp. nov. can be distinguished from all known species of Pantolyta by the combination of the following characters: the head is not nasiform (Fig. 8B) and has no mouth conus ( Fig. 2C); the genae are weakly convex in frontal view ( Fig. 2C); the female antenna is slender, weakly broadened apically, A15 is wider than A 14 in dorsal view ( Fig. 4C, E); A 3 in male has a deep emargination and a keel reaching mid-length of the segment ( Fig. 6D); the pronotal pit is bare, with a distinct vertical keel inside (Fig. 9D); the mesopleuron possesses subalar and epicnemial bridges and the mesopleural pit developed as a shallow depression (Fig. 10F); the propodeum has a simple median keel (Fig. 11B); upper and lower posterior propodeal projections form large teeth (Figs 11B and 13I); the fore wing has a closed radial cell, marginal vein longer than the distance from marginal vein to basal vein and almost as long as radial cell; the petiole is transverse.

Etymology. This new species is named in honour of the entomologist Igor Melnik (Moscow), who collected the holotype of this species.

Distribution. Russia (Kunashir Island), Republic of Korea.

Hosts. Unknown.