Pantolyta dichromia, Chemyreva & Kolyada, 2021

Pantolyta dichromia sp. nov.

( Figs 1B, 4B, 6C, 8A, 10D, 11F, 13J, 14A)

Holotype. Female; Russia, Primorskiy Terr., Lazovskiy Nature Reserve , 43°15′17″N 134°07′59″E, yellow pan trap, 5–25.VII.2005, K. Makarov leg. ( ZIN). GoogleMaps

Paratypes (all in ZIN). Russia: Samara Prov.: Stavropol'skiy Distr., Zhiguli Nature Reserve, near Bakhilova Polyana , 3.VII.2011, Chemyreva leg., 1 female ; Kinel’ Distr., near Domashka, forest along Samara River , 29–31.V.2019, 16–18.VI.2019 and 2–4. VII.2019, Chemyreva leg., 1 male, 4 females ; Primorskiy Terr. : same locality and data as for holotype, 2 females ; same locality as for holotype, VII.2007, K. Makarov leg., 3 females GoogleMaps ; same locality as for holotype, 9–17.VII.2008, A. Khalaim leg., 1 male GoogleMaps ; Sakhalin Prov., Kunashir I., vicinity of Sernovodsk , 15–16. VII.1973, D. Kasparyan leg., 1 male . Azerbaijan, Lenkoran’ , 3–7. V.1971, V. Tobias leg., 1 male .

Description. Female (holotype). Body length 3.4 mm; fore wing length 2.3 mm; antenna length 1.9 mm.

Coloration. Head, A13–A15, mesosoma and petiole dark brown; T2–T6 and S2–S5 brown; A1–A12, mandibles, veins and metasoma (excluding petiole) pale brown; palpi, legs, T7–T8 and S6 yellowish brown.

Head not nasiform in dorsal view, 0.77 times as long as wide, wider than mesosoma, smooth, with few scattered long setae. Temples sharply receding behind eye in dorsal view. Occipital flange narrow, sculptured. Head in lateral view 1.26 times as high as long. Malar space 0.71 times as long as largest diameter of eye. Eye weakly pubescent. Antennal shelf in frontal view with fine coriaceous sculpture below toruli. Face smooth, with long scattered setae. Head in frontal view with mouth conus indistinct; genae converging towards mouthparts in frontal view. Clypeus transverse, 0.67 times as high as wide. Tentorial pits small. Labrum small, semicircular, hardly discernible. Mandibles asymmetrical ( Fig. 1B).

Antenna slender, weakly broadened apically ( Fig. 4B). Ratios of length to width of A1–A 15 in dorsal view: A1 (70: 12); A2 (20: 10); A3 (24: 9); A4 (18: 9); A5 (16: 10); A6 (16: 10); A7 (16: 10); A8 (15: 10); A9 (15: 10); A10 (15: 12); A11 (15: 12); A12 (14: 12); A13 (14: 13); A14 (15: 14); A15 (34: 16).

Mesosoma slightly compressed, 1.1 times as high as wide, in dorsal view 1.8 times as long as wide. Sides of pronotum bare, pronotal collar with scattered long setae; epomia strongly prominent; pronotal pit bare, with epomia inside. Mesoscutum 1.19 times as long as wide, convex, with few scattered semierect setae. Notauli deep and complete, not sculptured. Anterior scutellar pit deep and rounded. Scutellum large, widened posteriorly. Axillar depression scarcely pubescent, with verriculate tubercle anteriorly (Fig. 11F). Mesopleuron scarcely pubescent ventrally and anteriorly, with subalar bridge posterodorsally and epicnemial carina anteroventrally; epicnemial pit deep and pubescent; mesopleural pit completely absent (Fig. 10D). Metascutellum narrow, pubescent; dorsellum distinct, with three short longitudinal keels. Propodeum transverse, 0.68 times as long as wide, bare dorsally and pubescent laterally, medi- an propodeal keel bifurcate from base; upper and lower posterior propodeal projections forming distinct teeth (Figs 10D and 11F). Sides of propodeum entirely pubescent, with three longitudinal keels (Fig. 10D). Legs slender.

Fore wing with closed radial cell (Fig. 14A). Distance from marginal vein to basal vein / marginal vein length / radial cell length as 15 / 16 / 16.

Petiole of metasoma cylindrical, as long as wide, with coarse rugose sculpture, longitudinal keels and several rows of long erect setae laterally and ventrally; petiole ventrally with several bunches of setae; posterior margin of petiole weakly arcuate (Fig. 11F). T2 anteriorly with short striation, median groove the longest; T3–T6 narrow, with few setae; T7 and T8 long, strongly compressed laterally. S2 with short grooves at base and with numerous erect scattered setae; S3–S5 narrow, with scattered setae; S6 distinctly elongate and compressed, with a few scattered setae.

Male. Body length 2.9–3.2 mm. Similar to female except following characters: antenna filiform, slightly longer than body; antennomeres cylindrical, with uniform short pubescence; A1 slender, 0.55 times as long as head width; A3 deeply emarginate, with keel extending to one-third of A3 length ( Fig. 6C); ratios of length to width of A1–A 5 in lateral view: A1 (50: 12); A2 (14: 11); A3 (50: 12); A4 (44: 11); A5 (44: 11); radial cell longer than marginal vein; petiole elongate, 1.57 as long as wide (Fig. 13J); T7 transverse, not compressed; S2–S7 with numerous semierect scattered setae; T8 and S8 subtriangular, rounded apically.

Variation. Female. Body length 3.2–3.7 mm. Median propodeal keel bifurcate from base to posterior 0.65; branches of median propodeal keel adjacent to each other or distinctly separated. Distance from marginal vein to basal vein, length of marginal vein and length of radial cell equal to each other, or sometimes marginal vein longer. Petiole as long as wide or slightly elongate.

Comparison. Pantolyta dichromia sp. nov. can be distinguished from all known species of Pantolyta by the combination of the following characters: the head is not nasiform (Fig. 8A) and has no mouth conus, the genae converge toward mouthparts ( Fig. 1B); the female antennae are slender, weakly broadened apically, A3–A10 are yellowish, with two or three apical segments dark brown ( Fig. 4B);

V.G. Chemyreva & V.A. Kolyada. Taxonomy of Acropiesta , Anommatium , Erasikea and Pantolyta

A3 is deeply emarginated, with a keel extending to one-third of A3 length ( Fig. 6C); pronotal pit is bare, with short epomia inside; the mesopleuron possesses subalar and epicnemial bridges, the mesopleural pit is completely absent (Fig. 10D); the median keel is bifurcated from the base; upper and lower posterior propodeal projections form large teeth (Figs 10D and 11F); the fore wing has a closed radial cell.

Etymology. The name derives from the Ancient Greek adjective “δί- χρωµος” (bicolor), in reference to the bicolourous antennae of female.

Distribution. Azerbaijan, Russia (European part, Primorskiy Territory, Kunashir Island).

Hosts. Unknown.