Lathyrus digitatus, (Bieb.) Fiori Lathyrus pannonicus, (Jacq.) Garcke Lathyrus pannonicus subsp. pannonicus Lathyrus pannonicus subsp. collinus, (Ortmann) Soo Lathyrus pannonicus subsp. asphodeloides, (Gouan) Bassler

14. L. digitatus (Bieb.) Fiori View in CoL in Fiori & Paol., FI. Anal. Ital. 2: 105 (1900)

(incl. L. sessilifolius (Sibth. & Sm.)Ten. , L. tempskyanus (Freyn & Sint.) K. Malÿ ).

Glabrous perennial; stem 10-40cm, not winged; roots fusiform, fleshy. Leaves usually almost digitate, without tendril; leaflets 1-2 pairs, 15-80x2-8 mm, linear; stipules 6-8 x 0-5-1 mm, lanceolate, semi-sagittate. Racemes 4- to 10-flowered. Calyx-teeth slightly unequal; corolla 15-30 mm, bright reddish-purple. Legume 40-70 x 5-9 mm, brown, glabrous. Seeds 5-7, smooth; hilum f of the circumference. 2m = 14. S.E. Europe, S. Italy. Al Bu G r It Ju Rs (K) Tu.

15. L. pannonicus (Jacq.) Garcke View in CoL , Fl. Nord-Mittel-Deutschl. ed. 6, 112 (1863)

(L. albus (L. fil.) Kittel ).

Glabrous or sparsely pubescent perennial; stem 15-50 cm, not or very narrowly winged; roots tuberous, fleshy. Leaves without tendril; leaflets 1-4 pairs, 15-75 x 2-5(-8) mm, linear to oblong-lanceolate or oblong-elliptical; stipules 10-20x 1-2-5 mm, linear-lanceolate, semi-sagittate. Racemes 3- to 9-flowered. Calyx glabrous or ciliate on the margin, the teeth unequal; corolla 12-20 mm, pale cream with reddish cr purplish tinge; style filiform. Legume 30-65x3-8 mm, pale brown, glabrous. Seeds 12-20, smooth; hilum I of the circumference. 2n= 14. From C. Spain, S. Italy and S.E. Russia northwards to N. W. France and to 54° N. in C. Russia. Al Au Bu Cz Ga Ge Hs Hu It Ju Po Rm Rs (C, W, K, E). A variable species that has been variously divided into varieties and subspecies or even separate species. There is no general agreement as to the number of taxa that can be recognized. The fullest account is given by Sirjaev, Bull. Assoc. Russe Sci. Prague (Sci. Nat. Math.) 5: 239-261 (1937), who recognized 6 varieties in Europe. Bässler, Feddes Repert. 72: 89 (1966), gives a list of 6 subspecies from Europe, without any explanation, which differs somewhat from Sirjaev’s treatment, a modification of which is given here. 1 Lowest calyx-tooth about as long as or longer than tube; stem usually very narrowly winged near the apex (e) subsp. varius 1 Lowest calyx-tooth shorter than tube (sometimes only slightly shorter); stem not winged

2Leaflets 15-30 x 4-7 mm, oblong-lanceolate or oblongelliptical (d) subsp. hispanicus 2 Leaflets 35-75 x 2-6(-7) mm, linear or linear-lanceolate or linear-elliptical

3 Peduncles shorter than or about as long as the subtending leaf; calyx-teeth usually glabrous, the lowest as long as the tube (b) subsp. collinus 3 Peduncles usually exceeding the subtending leaf; calyx-teeth usually ciliate, the lowest usually not more than | as long as the tube

4 Tubers 2-5 cm, oblong-ovoid (a) subsp. pannonicus 4 Tubers mostly 10 cm or more, slender or fusiform

(c) subsp. asphodeloides

(a) Subsp. pannonicus :

Stems up to 50 cm, not winged; tubers 2-5 cm, oblong-ovoid. Leaflets 35-60 x 2-4 mm, linear or linearlanceolate. Peduncles 6-9 cm, usually exceeding the subtending leaf. Calyx-teeth ciliate, the lowest c. i as long as tube. Damp pastures. E.C. Europe and N. W. part of Balkan peninsula.

(b) Subsp. collinus (Ortmann) Soó View in CoL , Scripta Bot. Mus. Transs. 1: 46 (1942)

( L. lacteus (Bieb.) Wissjul. , L. pannonicus var. versicolor auct. pro parte):

Stems up to 50 cm, not winged; tubers usually more than 10 cm, slender or fusiform. Leaflets 35-70 x 2-6(-7) mm, linear, linear-lanceolate or linear-elliptical. Peduncles 3-7(-8) cm, usually shorter than the subtending leaf. Calyxteeth usually glabrous, the lowest i - f as long as tube. Dry grassland and scrub. E.C. & E. Europe, N. part of Balkan peninsula.

(c) Subsp. asphodeloides (Gouan) Bässler View in CoL , Feddes Repert. 72: 89 (1966):

Like subsp. (b) but peduncles 5-9 cm, usually exceeding the subtending leaf; calyx-teeth usually ciliate, the lowest c. i as long as tube. Dry grassland and scrub. From W. France to the Alps and S. Italy.

(d) Subsp. hispanicus (Lacaita) Bässler View in CoL , loc. cit. (1966):

Stems 15-30 cm, not winged; tubers usually more than 10 cm, slender or fusiform. Leaflets 15-30x4-7 mm, oblong-lanceolate or oblong-elliptical. Peduncles 4-8 cm, usually exceeding the subtending leaf. Calyx-teeth glabrous or sparsely ciliate, the lowest c. i as long as tube. Dry grassland and scrub. • C. & E. Spain.

Intermediates between subspp. (c) and (d) occur in the Pyrenees and in N.E. Spain. This subspecies has also been recorded from Krym but all the specimens seen from there are referable to subsp. (b).

(e) Subsp. varius (C. Koch) P. W. Ball View in CoL , Feddes Repert. 79: 47 (1968)

( L. pannonicus var. versicolor auct. pro parte):

Stems up to 50 cm, narrowly winged, at least near the apex; tubers 3 cm or more, oblong-ovoid to fusiform. Leaflets 35-75 x 3-7 mm, linearlanceolate or linear-elliptical. Peduncles 5-11 cm, as long as to much longer than the subtending leaf. Calyx-teeth usually ciliate, the lowest as long as or longer than tube. Dry grassland and scrub. • N. Italy, N. W. Jugoslavia.

Very variable in the size of the tubers and the length of the peduncle. Plants with short tubers and short peduncles are possibly intermediate between subspp. (a) and (e).