Rubus ulmifolius, Schott

38. R. ulmifolius Schott View in CoL , Z s/'j 1818: 821 (1818)

( R. rusticanus Merc. , R. discolor sensu Syme , non Weihe & Nees, R. amoenus Portenschl. , non Koehler).

Stems robust, arching or procumbent, angled, often sulcate, pruinose, glabrous to tomentose with semiappressed hairs; prickles robust, broad-based, patent to falcate, hairy. Leaves pedate, often very small; leaflets 3-5, dark green and glabrous above, white-tomentose beneath, convex, variously toothed, coriaceous; terminal leaflet ovate or suborbicular to obovate. Inflorescence often long and narrow, sometimes pyramidal, leafy at the base, with patent branches and long pedicels; all axes with robust, broad-based, patent to falcate, hairy prickles. Sepals deflexed after flowering, sometimes slightly aciculate, white-tomentose; petals crumpled, orbicular or ovate, sometimes jagged at the apex, pink or occasionally white; stamens white or pink, equalling or just exceeding the green, pink or white styles; anthers glabrous; pollen completely fertile; carpels hairy, often tomentose. 2/2= 14. S., W. & C. Europe. Al Au Az Be BI Br Co Cr Cz G a Ge Hb He Ho Hs It Ju Lu Sa Si Tu.

The only diploid, sexual species in Subgen. Rubus whose chromosome number has been verified from material of wild origin; it is very polymorphic in shape of leaves, branching of inflorescence, clothing of the axis, size of flowers and colour of petals, and up to 20 subspecies as well as 92 varieties have been described.

Related species include: R. heteromorphus Ripart ex Genev. , Mém. Soc. Acad. (Angers) 24: 255 (1868) ( R. dalmatinus Tratt. ex Focke ). Al Ju. R. portuensis Samp. , Ann. Sci. Nat. (Porto) 8:10(1904). Hs Lu. R. sanguineus Friv. , Flora (Regensb.) 18: 334 (1835) ( R. sanctus auct. plur., non Schreber, R. anatolicus Focke , R. discolor Boiss. , non Weihe & Nees). Bu Rs (K). R. thessalus Halâcsy , Consp. Fl. Graec. 1: 503 (1900) (R.

anatolicus var. cinereus Hausskn. ). Al Gr.