Rosa sicula, Tratt. Rosa glutinosa, Sibth. & Sm. Agrimonia L. Agrimonia pilosa, Ledeb. Agrimonia eupatoria, L.

44. R. sicula Tratt. View in CoL , Rosac. Monogr. 2: 86 (1823)

( R. thuretii (Burnat & Gremii) Burnat & Gremii ).

Up to 0-5 m. Prickles usually sparse, slender, more or less curved, sometimes mixed with acicles and glandular setae. Leaflets 5-7, 7-15 x 5-15 mm, broadly ovate to suborbicular, compound-serrate, usually glabrous and sparsely glandular above, densely glandular and glabrous or sparsely pubescent beneath. Pedicels 3-5 mm, often stipitate-glandular, sometimes sparsely pubescent. Sepals glandular on the back, patent or erect after anthesis. Petals 10-15 mm, pink. Styles villous to lanate. Fruit c. 10 mm, globose, usually sparsely stipitate-glandular, red. Mediterranean region. Al Ga G r Hs It?Ju Si.

45. R. glutinosa Sibth. & Sm. View in CoL , FI. Graec. Prodr. 1: 348 (1809).

Like 44 but prickles mixed with numerous stalked glands and setae; leaflets more densely glandular and usually densely pubescent to tomentose on the upper surface. E. & C. Mediterranean region, Balkan peninsula. Al Bu Cr G r It Ju Si.

46. R. turcica Rouy View in CoL , III. Pl. Eur. Rar. 6: 45 (1896)

( R. ferox Bieb. , non Lawrance, R. horrida Fischer ex Crépin , non Sprengel).

Like 44 but prickles abundant, stout, curved, mixed with numerous setae and stalked glands; sepals deflexed and deciduous after anthesis; styles glabrous or somewhat villous. S.E. Europe. Bu G r Rm Rs (W, K) Tu.

47. R. serafinii Viv. View in CoL , FI. Lib. 67 (1824).

Like 44 but prickles hooked or falcate, rarely mixed with setae; leaflets 8-12 x 6-10 mm, shining and glabrous above; pedicels glabrous; sepals glandular or eglandular, deflexed and deciduous after anthesis; fruit glabrous. C. Mediterranean region; Bulgaria and S. Jugoslavia. Bu Co It?Rm Sa Si.

11. Agrimonia L. 1

Perennial, rhizomatous herbs. Stem erect, with glandular hairs. Leaves irregularly pinnate. Flowers 5-merous, in terminal, spikelike racemes; pedicels short, with2bracteoles. Hypanthium deeply concave, becoming hard in fruit; epicalyx absent; stamens 5-20; carpels 2; style terminal; ovules 1. Fruit of 1 to 2 achenes enclosed in the hypanthium, which is obconical, turbinate or cylindrical, and has hooked bristles at the upper end.

Literature: V. Skalicky, Acta Horti Bot. Prag. 1962: 87-108 (1962).

1 Petals pale yellow; mature hypanthium (including bristles) 4-5 mm 1. pilosa 1 Petals golden yellow; mature hypanthium (including bristles)

6-12 mm

2 Pedicels 4-10 mm; bracts often ovate and entire 4. repens

2 Pedicels 1-4 mm; lower bracts trilobed

3 Stem with both short and long eglandular hairs; mature hypanthium grooved for at least J of its length 2. eupatoria

3 Stem with long eglandular hairs only; mature hypanthium grooved for half its length 3. procera

1. A. pilosa Ledeb. View in CoL , Ind. Sem. Horti Dorpat., Suppl. 1 (1823)

( A. dahurica Willd. ex Ser. ).

Stem 50-150 cm, with long eglandular hairs. Leavps green on both surfaces, with glandular but very few eglandular hairs beneath; leaflets with 3-8 pairs of teeth, cuneate and entire at base. Raceme lax. Petals 2-5-4 mm, pale yellow, entire. Mature hypanthium (including bristles) 4-5 x 3-3-5 mm, deeply grooved throughout its length, more or less glabrous except for the bristles; inner bristles connivent and interwoven. U.S.S.R. from c. 49° to c. 62° N., extending westwards to S. Finland, N.E. Poland and C. Romania. Fe Po Rm Rs (N, B, C, W) [Cz].

2. A. eupatoria L. View in CoL , Sp. Pl. 448 (1753) View Cited Treatment .

Stem 15-150 cm, with both long patent, and short flexuous, eglandular hairs. Basal leaves often in a rosette and basal internodes short. Leaves with 3-6 pairs of main leaflets and 2-3 smaller pairs in between; leaflets serrate or crenulate almost to the base, dark green above, whitish- or greyish-tomentose beneath, with glandular hairs; stomata (21-)23-25(-27) p. Pedicels at maturity 1-3 mm; lower bracts trilobed. Petals (3-)4-5(-6) mm, golden yellow, obovate, usually not emarginate. Mature hypanthium obconical to turbinate, deeply and narrowly grooved for at least | of its length, and with many eglandular, appressed hairs; the inner bristles erect, the outer ascending, patent or deflexed. Almost throughout Europe except the extreme north. All except Cr Fa Is Sb.

(a) Subsp. eupatoria :

Plants with rosette 15^40 cm, those without rosette up to 150 cm; not villous, except occasionally the tallest plants. Leaflets broadly obovate and coarsely crenulate to elliptical and serrate; glandular hairs concealed by the tomentum. Hypanthium (including bristles) 7-1 0x 5-7 mm, grooved throughout its length; disc scarcely projecting; lowest bristles ascending or patent. 2n = 28. Throughout the range of the species, except Açores.

(b) Subsp. grandis (Andrz. ex Ascherson & Graebner) Bornm. View in CoL , Feddes Repert. (Beih.) 89:244 (1940):

Plants up to 150 cm, robust, villous, usually without rosette. Leaflets elliptical, coarsely serrate; glandular hairs concealed by the tomentum. Hypanthium (including bristles) 1 1x9 mm, grooved throughout its length; disc projecting by c. 0-4 mm; lowest bristles patent or slightly deflexed. S., E. & E.C. Europe.

(c) Subsp. asiatica (Juz.) Skalicky View in CoL , Feddes Repert. 79: 35 (1968)

( A. asiatica Juz. ).

Without rosette. Leaflets elliptical, coarsely and acutely dentate; glandular hairs not concealed by the tomentum. Hypanthium (including bristles) 7-8 x 7-8 mm, grooved for I of its length; disc projecting by c. 0-5 mm; lowest bristles always distinctly deflexed. S.E. Russia. (C. & S.W. Asia.)

3. A. procera Wallr. View in CoL , Erst. Beitr. FI. Hercyn. 203 (1840)

( A. odorata auct., non Miller).

Stem 50-120 cm, with long eglandular hairs; without basal rosette. Leaves green on both surfaces; stomata (27-)30(-33) leaflets elliptical, coarsely and acutely serrate almost to the base. Pedicels at maturity (1 -5—)2-5(—4) mm; lower bracts trilobed. Petals golden yellow, oblong to obovate, often emarginate. Mature hypanthium (including bristles) c. 11 x II mm, turbinate, with broad, shallow grooves for c. $ its length, and with well-developed disc; lowest bristles distinctly deflexed.

2n = 56. W., C. & S. Europe, extending to S. W. Finland and E. Ukraine. Au Be Br Bu Cz Da Fe Ga Ge Hb He Ho Hs Hu It Ju Lu No Po Rm Rs (B, C, W) Si Su.

4. A. repens L. View in CoL , Syst. Nat. ed. 10, 2: 1046 (1759)

( A. odorata Miller ).

Stem 50-100 cm, villous, with long and short eglandular hairs; without basal rosette. Leaves large, coriaceous, dark green above, greyish-green beneath; leaflets overlapping. Stipules large. Pedicels at maturity (4-)5(-10) mm; bracts often ovate and entire. Petals 5-7 mm, golden yellow, oblong, rounded at apex. Hypanthium (including bristles), 10-12 x 12-14 mm, cylindrical, with deep and narrow grooves for | of its length; disc stout, projecting by c. 1 -5 mm; lowest bristles distinctly deflexed. Formerly cultivated in gardens and now naturalized in various parts of Europe. [Au Be Cz Ge?It Rm Tu.] (Anatolia.)