Turgaphagites Lyubarsky et Perkovsky, 2024

Tribe Cryptophagini Kirby, 1826

Turgaphagites Lyubarsky et Perkovsky gen.nov.

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Figs 1–2 View Figure 1 View Figure 2

Type-species: Cryptophagites clavatus Ponomarenko, 1990

Description. Body small ( 2.7 mm), moderately convex, parallel-sided ( Figs 1A, B View Figure 1 ). Head prognathous, constricted behind. Antennal insertions exposed in dorsal view, inserted into small concavity, antennae relatively short, not reaching hind margin of pronotum, with three-segmented club. Antennae 11- segmented. Flagellomeres elongate; segments of club transverse. Eyes large, about half length of head. Lateral edge of pronotum without callosity, teeth, and serrations. Elytra oval, cover entire abdomen, not truncate. Pygidium not exposed. Prosternum in front of procoxae short. Prosternal process narrow, not extended beyond or only slightly reaching beyond posterior edge of procoxae, rounded apically. Procoxal cavity half-open internally. Mesocoxal cavities laterally closed (?) by meeting of mesoventrite and metaventrite. Shortest distance between mesocoxal and metacoxal cavities distinctly less than half as great but less than shortest diameter of cavity. Metathorax short, its length 0.4 times of width, significantly longer than mesothorax ( Figure 2 View Figure 2 ). Metaventrite shorter than prothorax and mesoventrite combined. Metacoxae slightly transverse, shortest distance between metacoxal cavities less than longest diameter of cavity. Epipleuron distinct in basal half, reduced in the posterior half of elytron. Hind wings present. Legs slender. Procoxae small, rounded, coxae close together. Distance between them approximately equal to diameter of coxae. Tibia without strong teeth. Metatarsi 5-segmented (tarsal formula?–5–5), segments not lobed. Tarsomeres elongated. Last tarsomere longest, its length 0.7 times of length other tarsomeres together. Abdomen with five visible segments. Abdominal intercoxal process broadly rounded. Abdominal ventrite 1 longer than 2.

Unfortunately, the dorsal aspect is not visible. We assume that the punctae of the elytra are confused.

Remarks. We assign Turgaphagites gen. nov. to the Cryptophagidae , as it corresponds to the diagnosis of the family even without observing confused punctures of the elytra. As the tibiae are slender, not club-shaped, this genus probably belongs to the tribe Cryptophagini .

Etymology. The genus name Turgaphagites is a combination of Turga ( type locality) and Cryptophagites Ponomarenko, 1990 .

Comparison. The new genus differs from other Cretaceous genera of Cryptophagidae by the terminal tarsomere of hind leg being shorter, its length 0.7 times the length of the other tarsomeres together, and the metasternum being short, 0.4 times as long as wide. The new genus differs from extant genera Micrambe and Mnioticus by short metasternum.

Turgaphagites clavatus ( Ponomarenko, 1990) , comb. nov.

Figs 1–2 View Figure 1 View Figure 2

Ponomarenko & Ryvkin 1990: 78 ( Cryptophagites )

Material. Holotype PIN 1742 View Materials /169 Turga , Turga Formation, Lower Cretaceous.

Description. Length 2.7 mm, width 1.2 mm, length of elytron 1.75 mm. Body moderately convex; parallel-sided ( Figure 1 View Figure 1 ). Head constricted behind eye. Eye well-developed. Temples approximately equal to length of eye. Subgenal spine not developed. Gular sutures spaced. Antennae relatively short, not reaching pronotum hind margin, inserted in small concavity, with 3-segmented club. Scape large, much wider than the other flagellum segments. Pedicel equal in length to scape, but significantly narrower. Antennomeres 3, 4, 6, 7 elongate, 8 subquadrate. Antennomeres 9 and 10 transverse, 11 elongated, not narrower than 10. Club 0.55 times as long as other antennomeres combined. Pronotum, transverse, 1.5 times as wide as long, greatest width at anterior third, slightly narrowed anteriorly, strongly narrowed posteriorly, margin unmodified, angularity absent. Base of pronotum noticeably narrower than elytra at humeri. Elytra elongated, 2.9 times longer than pronotum, rounded apically, widest in anterior third. Width of prosternal process less than procoxa. Prosternal process rounded apically. Width of mesoventral process less than mesocoxa. Metathorax 1.9 times as long as mesothorax. Abdominal ventrite 1 without (?) subcoxal line; intercoxal process about as long as wide. Abdominal segments length ratio 1.1: 1: 1: 1: 1.2. Ventrite 5 covered by confused punctation, posterior margin without crenulations. Tibiae slender, weakly dilated apically, without large tooth or spine. Tarsomeres 1–3 of protarsus not dilated. Tarsomeres 1–5 of metatarsus elongated, ratios 1.3: 1.3: 1: 1: 1.8 ( Figure 2 View Figure 2 ).