Elwendia bucharica Kljuykov & Lyskov, 2018

Elwendia bucharica Kljuykov & Lyskov View in CoL sp. nov. ( Fig. 1 View FIGURE 1 , 3a View FIGURE 3 , 4a View FIGURE 4 , Table 1)

The new species differs from E. capusii in shorter terminal leaf lobes ( 1–4 mm vs. 10–40 mm), bracteoles (wanting or 1–3 herbaceous bracteoles vs. (3) 5–8 bracteoles with membranous margins), pedicels ( 2–3 mm long, ribbed, slightly thickened vs. up to 5 mm long, tiny, terete), and the number of secretory ducts in petals (several vs. solitary).

Type: — UZBEKISTAN. Southern side of Hissar Range, near Xojaosmin, upper reaches of Xondiza River ( 38.626° N, 67.606° E), 07 May 1979, Pimenov, Kljuykov, Boryaev, Baranova et Vasilyeva 483 ( holotype MW 0595669, isotype MW 0595670).

Polycarpic herbaceous plant with tuberous root, all glabrous; tuberous root obovate, up to 2 cm ⌀. Stems 10–15 cm tall and about 1–2 mm ⌀ at base, solitary, solid, angular-ribbed in lower part, tiny-ribbed under umbels, bifurcated from lower stem part, lower internodes short. Radical leaves petiolate; leaf blades 2–3.5 cm long, 2.5–3 cm wide, triangular or rhombic, 3–4-ternate; basal segments with petiolulate 7–9 mm long, terminal segments lanceolate or obovate, deeply dissected into linear or lanceolate 1–4 mm long lobes; lower stem leaves sessile or with short, utricular, lanceolate or linear-lanceolate sheaths; 2–3-pinnate, leaf blades smaller than radical leaves; upper stem leaves with narrow sheaths, leaf blades pinnate, terminal lobes filiform, 3–4 mm long. Umbels 4–5 cm ⌀, with 3–4 tiny, ribbed, equal rays up to 3.5 cm long, bracts wanting. Central umbel located at same level as terminal umbels of lateral branches. Umbellets 7–10 mm ⌀, up to 8–10 flowers, pedicels slightly unequal, thickened, ribbed, up to 3 mm long; bracteoles wanting or 1–3 narrowly linear, entire; calyx teeth obscure, petals white, 1.2–1.5 mm long, obovate, with narrow notch, elongate narrow curved apex, and several secretory ducts. Fruits nearly cylindrical ( Fig. 3a View FIGURE 3 , 4a View FIGURE 4 ); mericarps homomorphic, terete, 3–4 × 0.8–1 mm; only primary ribs present; all ribs equal, filiform, straight, with entire margin; stylopods low conical; styles 1–1.2 mm long, suberect; commissure broad; vascular bundles thin; secretory ducts single between ribs; two secretory ducts on commissural side; endosperm flat at commissural side.

Additional specimens examined ( paratypes): — UZBEKISTAN. Hissar Range , valley of Sangardak River, upper reaches of Oqsuv River ( 38.650° N, 67.383° E), juniper community, 5 June 1948, Pyataeva 208 ( TASH) GoogleMaps ; Toʻpolon River basin, Maland pass ( 38.618° N, 67.654° E), 17 June 1948, Pyataeva 471 ( TASH) GoogleMaps .

Etymology: —The specific epithet refers to the Khanate of Bukhara, a Central Asian state that existed from the second quarter of the 16th century to the late 18th century and included the territory from which the plants were collected.

Phenology: —Flowering in April or May; fruiting in June.

Distribution area: — Uzbekistan, Hissar Range, Sangardak River basin ( Fig. 5a, b View FIGURE 5 ).

Habitats: —The species grows on shallow gravel slopes among Prangos pabularia , at elevations of about 2000 m.