Glyptosmiini Engel, 2024

Glyptosmiini Engel , new tribe

TYPE GENUS: Glyptosmia Engel , new genus.

DIAGNOSIS: This tribe possesses a unique intermingling of features of Glyptapini , Dioxyini View in CoL , and Ctenoplectrellini as well as its own distinctive apomorphies. Like Glyptapini and Ctenoplectrellini the metatibia bears a scopa composed of abundant, long setae (fig. 13), although in this tribe many of the setae on the posterior surface are characteristically hooked apically, and some of those on the anterior surface have short apical branches. Likewise, the pretarsal claws have a short, subapical ramus, arolia on all legs, the basal vein is arched, and 2m-cu is slightly basal to 2rsm, and thus both 1m-cu and 2m-cu enter the second submarginal cell. Like Glyptapini the basal area and lateral surface of the propodeum are areolate and the mandible is long, slender, and has a large edentate margin above the apical tooth. Unlike glyptapines the compound eyes are bare, the omaular ridge is not carinate, and the mesosoma lacks large craterlike punctures and is instead covered with dense, strong, small punctures. Similar to Dioxyini View in CoL the metanotum is medially beset with tubercles, albeit paired in the fossil versus the single tubercle or spine in the former tribe. Unlike Dioxyini View in CoL , the sting and its associated structures are well developed. Quite distinctively, the mesoscutellum is flattened and projected posteriorly over the metanotum and the basal edge of the propodeum, and the apical margin is sinuate, and the metatibial spurs are noticeably thickened.

FEMALE. Integument without maculation and shiny black (fig. 13). Mandible with single apical tooth and edentate margin apical to expansion of trimmal extension; interdental laminae lacking. Malar space linear. Maxillary palpus tetramerous. Labial palpus tetramerous; palpomeres I and II flattened, elongate, palpomere I subequal to length of palpomere II; palpomere III projecting obliquely from axis of palpomere II. Clypeus somewhat flattened, not extending below lower tangent of compound eyes; clypeus apical margin not covering labral base, thus labroclypeal articulation not obscured. Single subantennal sulcus directed to outer margin of antennal torulus. Juxtantennal carina absent. Compound eyes bare (fig. 14). Posterior margin of vertex gently concave; preoccipital area sharply angled above but not carinate, otherwise rounded.

Pronotal collar virtually absent, posterior bor- der blending uninterrupted onto dorsal surface without transverse ridge medially to demarcate collar from dorsal surface of neck, with dorsal ridge rounded laterally without dorsolateral angle (carinate laterally in Glyptapini ), without dorsolateral ridge extending vertically across lateral surface. Mesoscutum broadly rounded anteriorly (fig. 13A, 15), raised above pronotal posterior margin, with dense, small punctures (without the deep craterlike punctures of Glyptapini ); parapsidal lines linear; mesoscutalmesoscutellar sulcus deeply impressed and wide (fig. 15), without V-shaped notch on mesoscutellum; mesoscutellum flat (figs. 15, 16A), posterior margin extending over metanotum and basal edge of propodeum (fig. 16A), apical margin with broad medial concavity (fig. 15); axillae simple (fig. 15); metanotum vertical, with pair of medial tubercles on either side of midline (fig. 16A); basal area of propodeum horizontal, asetose, areolate (as in Glyptapini ); omaular ridge angled but not carinate (carinate in Glyptapini ); scrobal sulcus absent; preepisternal sulcus absent; mesepisternum with dense, small punctures (areolate in Glyptapini ); dorsal lamella of metepisternum absent.

Forewing (fig. 17) with pterostigma proximal to r-rs longer than pterostigmal width; pterostigmal length more than 2× its basal width; prestigma more than 2× as long as broad; basal vein (1M) arched, thus orthogonal to 1Cu (as in Glyptapini and Ctenoplectrellini ); marginal cell apex rounded, slightly offset from anterior wing margin; two submarginal cells; 1m-cu and 2m-cu entering second submarginal cell, i.e., 2rs-m proximal to 2rs-m; hind wing with six distal hamuli.

Protibial calcar with anterior ridge on rachis bordering velum (as in Anthidiini View in CoL and Glyptapini ), malus simple, short, less than 0.5× length of velum, apical margin of velum simple; tibiae not spiculate; pro- and mesotibiae with apical outer spine, spine fainter on protibia; mesotibial spur long, slender, minutely serrate; metabasitibial plate absent; metatibial scopa present (figs. 13B, 18A) (as in Glyptapini and Ctenoplectrellini ; absent in other megachiline tribes); metatibial spurs long, distinctly thickened (fig. 18B), minutely serrate (not crescentic, ciliate, nor comblike); pretarsal claws with short inner ramus (fig. 18C, D); arolium present (fig. 18C).

Metasomal scopa present; sting and associated structures well developed (fig. 19).

MALE. Unknown.