Imeria (Caenojoppa) cephalotes ( ASHMEAD , 1905)

Imeria (Caenojoppa) cephalotes ( ASHMEAD, 1905) View in CoL (figs 4, 11, 23, 30, 37, 48)

Type not examined.

syn. nov. Elasmognathias albitarsis CUSHMANN, 1922

Type not examined.

S t u d i e d m a t e r i a l: Taiwan: Kankau (Koshun) Formosa, H. Sauter , 1♀ vii.1912 ; cephalotes det. Cushman ( SDEI); Cambodia: Siem Raep Prov., Angkor Thom , 1♀ 6-28.viii.2005, 1 ³ ix.2005, leg. D. R. Jump, Coll. I. R. Sc.N.B. (HK) ; Indonesia: N Sumatra, Aceh, Bengkung (Soraya), N.P. Gn. Leuser , Mal. trap, 1 ³ ii.1995, Y. van Nierop & Dolly (Leiden) ; Indonesia: Java, Verlaten Island [Sertung island between Java and Sumatra], 1 ³ xii. 1919, NHMUK 015214884 About NHMUK ( NHM) ; Pak Sha O, Hk , 50 Q KK 242849, 70 m, 1♀ 28.iii.-12.vi.2015, leg. C. Barth ( HK) ; 1♀ “M027.C.Hy.9” ( HK) ; 1♀ “M025.C.Hy.8” ( HK) .

D e s c r i p t i o n: ♀. Body length 8.5-9.5 mm. Flagellum with 35-40 segments, slender, bristle-shaped; 1 st flagellar segment 4.8-5.2× as long as wide, all segments much longer than wide. Temple very long, ± parallel behind eye, slightly and roundly narrowed apically. OED 1.8-2.0× and OOD 1.3-1.6× ocellar diameter. Temple and frons smooth and shining, with very sparse superficial punctures, frons medially with some superficial rugae. Face with sparse or rather dense punctures, shining. Gena with very sparse punctures. Clypeus almost flat, with very sparse punctures; apical margin sharp and pointed medially. Malar space 0.8× as long as width of mandibular base.

Epomia long but not reaching ventral margin of pronotum. Notaulus weakly impressed in frontal third of mesoscutum. Mesoscutum densely punctate, ± dull. Mesopleuron coarsely striate dorsally and rugose-punctate ventrally; speculum smooth. Epicnemial carina moderately widened behind fore coxa. Sternaulus weakly impressed in frontal 0.3 of mesopleuron. Metapleuron coarsely rugose-punctate; juxtacoxal carina absent. Scutellum moderately and roundly elevated, 1.1-1.3× as wide as long, rugose; lateral carina slightly widened, complete. Postscutellum ± striate, sometimes smooth. Propodeum rugose-punctate, completely carinated, without apophysis; spiracle long-oval. Area basalis c. 3× as wide as long, slightly bulging medially. Area superomedia hexagonal, about as long as wide; costula reaching at 4/10 of its length. Area petiolaris with transverse rugae. Hind coxa densely punctate, densely pilose, without or with weak scopa apico-ventrally. Hind femur with sparse punctures in ventral third, 4.2-4.3× as long as wide. Areolet pentagonal, frontal distance between veins 2rs-m and 3rs-m 2× their diameter. Vein 1cu-a interstitial.

Metasoma oxypygous. 2 nd to 4 th sternites with median folds. Postpetiole moderately widened, without latero-median carina, ± punctate laterally; median field not separated. Gastrocoelus impressed. Thyridium large, slightly oblique, c. 2× as wide as interval between thyridia. 2 nd tergite 1.2-1.4× as wide as long. 2 nd tergite densely rugose-punctate in basal half, densely punctate apically. 3 rd tergite with sparse or rather dense superficial punctures in basal half, almost smooth apically. Ovipositor sheath slender and stick-shaped, in profile 0.4-0.5 × as long as hind tibia.

Color: black. Mesosoma with pale setae. Flagellum black, flagellar segments 6-16/17 ivory dorsally. Palps, mandible except teeth, clypeus, face except brown median spot, spot on frontal orbit, gena and outer orbit, large spot on vertex separated from compound eye, frontal and upper margins of pronotum, two short paramedian stripes on mesoscutum, often tegula, subtegular ridge, wide longitudinal stripe on ventral mesopleuron, scutellum and postscutellum, apical spot on metapleuron, propodeal spots on area spiraculifera and on posterolateral areas, petiole mainly, often gastrocoelus and thyridium, sometimes narrow basal band on 3 rd tergite and apical bands on all tergites ivory. Fore and mid coxae and trochanters ivory; hind coxa black with ivory spots dorsally and laterally; fore and mid femora and tibiae reddish; fore and mid tarsi reddish or brown; hind femur red, often black basally in apical 0.2, sometimes with ivory spot apically; hind tibia red, black in basal 0.1-0.2 and apical 0.2-0.3; hind tarsus ivory, distal tarsomere usually black. Wings hyaline; pterostigma reddish-brown.

³. Body length 7.8-9.0 mm. Flagellum with 38-41 segments, bristle-shaped; 1 st flagellar segment 3.3-4.3× as long as wide, all segments longer than wide. Tyloids not visible. OED 1.4× and OOD 1.0× ocellar diameter. Face with fine sparse or rather dense punctures. Malar space 0.8-1.0× as long as width of mandibular base. Area basalis much wider than long. Area superomedia hexagonal, as long as or slightly longer than wide; costula reaching its middle. Hind femur densely punctate, 4.0-5.2× as long as wide. Postpetiole with fine microsculpture, finely punctate or partly with fine transverse rugae. 2 nd tergite 1.2-1.3× as wide as long, densely rugose-punctate. 3 rd tergite with dense superficial punctures. Following tergites shining and almost smooth. Apical margin of hypopygium with median tongue-like elongation which is almost pointed medially. Structure otherwise as in ♀.

Color: as in ♀. Flagellar segments 7/10-12/19 ivory dorsally. Palps, mandible except teeth, clypeus, face except brown median stripe, gena and outer orbit, frontal orbit, spot on vertex (separated from compound eye), frontal and upper margins of pronotum, short confluent paramedian stripes on mesoscutum, spot on tegula, subtegular ridge, wide stripe on ventral mesopleuron, scutellum, upper division and apical spot on metapleuron, area spiraculifera in front of spiracle, posterolateral stripes on propodeum, base of petiole, posterolateral spot on postpetiole, thyridium, apical bands on 2 nd to 7 th tergites and spot on paramere ivory. Fore and mid coxae and trochanters ivory; hind coxa with ivory dorsal and ventral spots; hind trochanter with ivory dorsal spot; fore and mid femora and tibiae blackish, with frontal ivory stripes; fore and mid tarsi ± black; hind femur reddish, black in basal 0.2 and apical 0.3, often with ivory apical spot; hind tibia reddish, basally and subapically blackish, apically ivory; hind tarsus entirely ivory or metasoma basally and distal tarsomere ± blackish. Wings hyaline; pterostigma black.

R e m a r k: In his original description of Elasmognathias albitarsis, CUSHMAN (1922) mentioned only minor differences to I. cephalotes (ASHMEAD) such as a distinctly elevated scutellum and slightly different color pattern of the hind tarsus. As already stated by HEINRICH (1934) in a longer series of I. cephalotes , the form of scutellum is variable and probably correlates with the size of the specimens. In my material, the color of hind tarsus is also variable. In recognition of this variability, both species are synonymized here.

D i s t r i b u t i o n: Known from the Philippines, Sulawesi and Taiwan, new records for Cambodia, Indonesia: Java and Sumatra, and continental China.