Imeria (Imeria) albomaculata formosana ( UCHIDA , 1930) Riedel & C, 2025

Imeria (Imeria) albomaculata formosana ( UCHIDA, 1930) View in CoL , stat. nov. ( figs 8 View Figs 5-10 , 20 View Figs 17-22 , 34, 39,

44) S t u d i e d m a t e r i a l: Taiwan: Kankau (Koshun), 3♀♀ 1 ³ iv.1912, 4♀♀ 2³³ vi.1912, 2³³

vii.1912, 2♀♀ viii.1912. 1 ³ ix.1912, 1 ³ x.1912, Taihorin, 1♀ iv.1910, 1♀ 1³ 7.xi.1911, 1³ 7.xii.1911, Hoozan, 1♀ 1³ 7.ix.1910, all leg. Sauter ( SDEI) ; Taiwan NW: Hueisuen exp. Forest , 600 m, 24°07´N 121°03´E, E Taichung, 1♀ 23-27.vi.2000, leg. W. Schacht ( ZSM) GoogleMaps ; Taiwan: (ROC), Hueisuen exp. Forest Stn (700 m), 24°05´N 121°02´E, 1♀ 3-8.ix.2002, leg. K. Schönitzer ( ZSM) GoogleMaps ; Taiwan: Pingtung, Chunri Township, Dahan Forest Road, N 24.41° E 120.72°, 2♀♀ 16.vi.2019, leg. K. Watanabe (Kanagawa) GoogleMaps .

D e s c r i p t i o n: ♀. Body length 8.5-9.5 mm. Flagellum with 38-40 segments, slightly lanceolate; 1 st flagellar segment 3.9-4.2× as long as wide, widest segments 1.3- 1.4× as wide as long. Temple long, slightly and roundly narrowed behind eye. OED 2.0× and OOD 0.9× ocellar diameter. Temple, frons and gena smooth and shining, with very sparse fine punctures. Face with dense superficial punctures. Clypeus large, flat, with fine very sparse punctures; apical margin slightly lamelliform raised laterally and with small tooth medially. Malar space 0.8× as long as width of mandibular base.

Epomia long, not reaching ventral margin of pronotum. Mesoscutum densely rugose, sometimes partly striate, dull. Notaulus not impressed. Epicnemial carina slightly widened behind fore coxa. Sternaulus impressed in frontal 0.4 of mesopleuron. Mesopleuron with coarse vermiform rugae, speculum ± smooth. Metapleuron rugose-punctate; juxtacoxal carina sometimes indicated at frontal margin of metapleuron. Scutellum almost pyramidal, 1.3× as wide as long, with complete lamelliform lateral carina. Postscutellum finely rugose. Propodeum incompletely carinated; spiracle long-oval; apical transverse carina weak but with posterolateral blunt tooth-like apophysis. Area basalis replaced by median longitudinal ridge. Area superomedia pentagonal, about as long as wide; costula reaching at 4/10 of its length. Lateral carina of area petiolaris and lateral longitudinal carina of propodeum weak. Hind coxa densely punctate, without scopa. Hind femur 4.5-4.7× as long as wide. Claws simple, apically strongly curved (by 90°). Areolet pointed frontally or pentagonal with frontal distance between veins 2rs-m and 3rs-m 1-2× their width. Vein 1cu-a slightly postfurcal by 1× its width.

Metasoma oxypygous, 2 nd to 5 th sternites with median folds. Postpetiole moderately widened, finely granulated but shining. Gastrocoelus moderately impressed, granulate. Thyridium large, transverse, c. 2× as wide as interval between thyridia. 2 nd tergite 0.9- 1.1× as long as wide. 2 nd tergite with dense superficial punctures in basal half, finely granulated apically. 3 rd tergite finely granulated. Ovipositor sheath widely reaching beyond metasomal apex, seen from lateral 0.3× as long as hind tibia.

Color: Black, metapleuron and propodeum usually ± reddish. Flagellum black, flagellar segment 6/7-14/15 ivory dorsally. Palps, mandible except teeth, clypeus ±, spots or stripes on facial and frontal orbits, sometimes spot on gena, frontal margin and hind edge of pronotum, tegula, subtegular ridge, sometimes postero-ventral spot of mesopleuron, scutellum, postscutellum, upper division of metapleuron, transversal band covering apical metapleuron, area petiolaris and area posteroexterna, base of petiole, wide basal bands of 2 nd and 3 rd tergites, narrow apical margins of 3 rd to 5 th tergites, apical band of 6 th tergite and 7 th tergite mainly or entirely ivory. Fore and mid coxae and trochanters ivory; hind coxa black or reddish, with dorsal ivory spot. Legs otherwise red; hind femur black in apical 0.1; hind tibia black in basal 0.1 and apical 0.1-0.2; hind tarsus reddish or brown. Wings hyaline; pterostigma reddish.

³. Body length 9-10 mm. Flagellum with 38 segments, bristle-shaped; 1 st flagellar segment 3.7-3.8× as long as wide. Tyloids on flagellar segments 9/10-17, linear to long-oval, maximally 0.5-0.6× as long as their flagellomeres. OED 2.0× and OOD 1.1× ocellar diameter. Mesopleuron striate dorsally and rugose-striate ventrally. Epicnemial carina moderately elevated behind fore coxa. Sternaulus impressed in frontal 0.3 of mesopleuron. Juxtacoxal carina sometimes partly present frontally. Spiracle long-oval to slit-shaped. Propodeum with small blunt apophysis. Hind femur 4.3-4.5× as long as wide. 2 nd tergite 1.1-1.2× as wide as long. Structure otherwise as described for the ♀.

Color: as described for the ♀ but propodeum sometimes mainly reddish; flagellar segments 8/9-15/18 ivory; facial orbit widely ivory, sometimes face entirely ivory; gena and outer orbit usually ivory; propleuron ± ivory, petiole ± ivory; paramere ivory. Hind tibia reddish, ± black basally and apically. Pterostigma reddish-brown.

R e m a r k: The Taiwanese specimens have only minor color differences compared to the Indian material of Imeria albomaculata . The propodeum and hind coxa are usually mainly red instead of black, but some specimens from the long series in SDEI have a completely black and ivory mesosoma. I was not able to find any other color or structural differences between the Indian and Taiwanese material. In my opinion, Imeria formosana (UCHIDA) represents a subspecies of I. albomaculata .

It should be noted here that the examined Imeria specimens from Vietnam, Laos and the Hainan Island of China belong to different but closely related species.

D i s t r i b u t i o n: Only known from Taiwan.