Empis (Enoplempis) undulasetosa, Sinclair & Brooks & Cumming, 2025

Empis (Enoplempis) undulasetosa sp. nov.

urn:lsid:zoobank.org:act:11CE9532-44A3-4A72-A016-26526BA11815

( Figs 230–232 View FIGURES 227–232 , 236 View FIGURES 233–236 , 238 View FIGURES 237–238 )

Type material. HOLOTYPE ♂, labelled: “USA: OR: Benton Co. / Corvallis ; Lewisburg Saddle / Old Growth trail; sweeping/ woods and creek/ 44.6423°N,- 123.2891°W / 23 April 2013, S. Fitzgerald ”; “ CNC/ 1078370 View Materials ”; “ HOLOTYPE / Empis (Enoplempis) / undulasetosa Sinclair ,/ Brooks & Cumming [red label]”( CNC) GoogleMaps . PARATYPES: CANADA. British Columbia: Agassiz [49°14′N 121°46′W], 4.iv.1925, R. Glendenning (2♂, CNC) GoogleMaps . USA. California: Humboldt County: Prairie Ck Redwoods SP, Drury Pkwy at Prairie Ck tr., 41.408°N 124.031°W, 3.vi.2009, E09-43b, B.M. Wiegmann et al. (1♂, CNC) GoogleMaps . Oregon: Benton County: Corvallis, Lewisburg Saddle , below Old Growth Tr. , 44°38′32.28″N 123°17′20.76″W, MT over creek, 23.iv–6.v.2013, SJF (9♂, 13♀, CNC) GoogleMaps ; same data except, 6–31.v.2013 (2 ♀, CNC) GoogleMaps ; 4 mi. up Woods Ck Rd from Hwy 20 [44.5440°N 123.5030°W], 9.v.2013, along Woods Ck, SJF (1♂, CNC) GoogleMaps ; same data except, 15.v.2014, sweep banks of Woods Ck (1♂, 1♀, CNC) GoogleMaps . Lane County: Oakridge , 18 mi. NW [43°52′N 122°42′W], 12.v.1969, E.M. & J.L. Fisher (1♂, CAS; 2♂, CSCA) GoogleMaps ; Siltcoos Lk., Camp Baker , 43°53′24″N 124°05′33″W, 54 ft, 10.v.2011, J.D. Pinto, R. L. Wescott (1♂, 1♀, UCRC) GoogleMaps ; same data except, 43°53′03″N 124°05′46″W, 20 ft (1♂, UCRC) GoogleMaps . Lincoln County: Waldport , 44°25′36″N 124°03′05″W, 56 m, 27–31.v.2009, MT, J.D. Pinto (1♂, UCRC) GoogleMaps ; same data except, 1–15.v.2010 (3♂, 3♀, UCRC) GoogleMaps ; same data except, iv.2011 (1♂, UCRC) GoogleMaps . Washington: King County: 5 mi. W Skykomish [47°42′N 121°21′W], 1.v.1977 (1♂, 1♀, WSU) GoogleMaps ; 10 mi. W Snoqualmie Pass [47°23′N 121°31′W], S. Fk Snoqualmie R, 15.vi.1975, N.E. Woodley (1♂, 3♀, WSU) GoogleMaps . Pacific County: Ft Canby SP nr Ilwaco [46°17′N 124°04′W], 13–15.vi.1973, WJT (1♀, WSU) GoogleMaps ; Ilwaco , 5.v.1918, ALM (2♂, USNM) ; same data except, 20.v.1918, A. Spurler (6♂, 1♀, WSU) ; South Bend [46°39′N 123°48′W], 23.v.1917, ALM (2♂, USNM) GoogleMaps . Whatcom County: Glacier [48°53′N 121°56′W], 4.vi.1917, H.G. Dyer (2♂, USNM) GoogleMaps .

Diagnosis. This species is distinguished from other species of the E. (En.) teres species group by the pair of dark scutal vittae, male hind femur with ventral knob-like process with tuft of whip-like setae ( Figs 231, 232 View FIGURES 227–232 ), and epandrium with long, wavy setae.

Description. Wing length 5.5–6.5 mm. Male. Similar to E. (En.) teres , except as follows: head with ocellar triangle with pair of long ocellar setae, longer than ocellar triangle.

Thorax with scutum with setae increasing in length posteriorly, with posterior seta as long as npl; 3 posterior npl. Scutellum with 1 pair of long sctl, with 1 pair of marginal setae.

Hind femur with white ventral pile on basal three-quarters; apical fourth slightly curved and swollen, with oval, knob-like ventral process bearing posterior appendix bearing whip-like tuft of black setae; tuft of anteroventral setae without swelling posterior to knob-like process; cluster of short, curved posteroventral setae posterior to knob-like process, with preapical oblique rows of 4–5 wavy posteroventral setae, as long as width of femur (not homologous to preapical row of posterior setae); without strong preapical anterior seta ( Figs 231, 232 View FIGURES 227–232 ). Hind tibia greatly narrowed at base; anteroventral digitiform process short, triangular with oblique row of appressed setae, as long as process; posteroventral process short, bifid with tufts of apical setae widely separated; ventral setae distal to processes not thickened; long anterodorsal and posterodorsal setae on apical two-thirds, more than twice as long as width of tibia ( Figs 230–232 View FIGURES 227–232 ). Hind tarsus with similar long dorsal setae.

Abdomen with sternites with long, wavy setae ( Fig. 230 View FIGURES 227–232 ). Terminalia ( Fig. 236 View FIGURES 233–236 ): brown with paler phallus. Cercus short, subtriangular, strongly tapered posteriorly to narrow apex; dorsal margin straight, with setae evenly distributed; setae shorter than half width of cercus; base fused to epandrial lamella. Subepandrial process extending to phallus. Epandrial lamella narrow, linear, longer than height; posterior margin strongly upturned; bearing long, wavy setae ventrally, longer than width of lamella. Hypandrium very short, truncate, not prolonged laterally; without setae. Phallus with basal third not greatly expanded, tapered, gently arched anteriorly; apex emerging beyond cercus; apex slender and downcurved in lateral view; with roughened, spiculate surface opposite dorsal margin of epandrium; ejaculatory apodeme oval, half-length of epandrium, Y-shaped, with ventrally positioned lateral apodemes.

Female. Similar to female of E. teres , except setae of scutum and scutellum longer.

Geographical distribution and seasonal occurrence ( Fig. 238 View FIGURES 237–238 ). Empis (En.) undulasetosa sp. nov. is known from southern British Columbia, Washington, Oregon and northern California, primarily along the Coast Range. Adults have been collected from April to June.

Etymology. The species name is Latin for wavy bristles, in reference to the long, wavy setae on the epandrium of this species.

Nuptial gift presentation. Form unknown.