Atanycolus Foerster, 1863

Genus Atanycolus Foerster, 1863 View in CoL View at ENA

Atanycolus Foerster, 1863: 238 View in CoL (name published in key, type species); Dalla Torre, 1898: 295 (catalogue); Ashmead, 1900: 138 (in key); Szépligeti, 1904: 18 (catalogue, distribution); Roman, 1910: 112 (diagnosis); Viereck, 1914: 16 (type species); Fahringer, 1926: 116 (diagnosis, key to species); Telenga, 1936: 26 (in key); Watanabe, 1937:13 (key to species of Japan, hosts); Shenefelt, 1943: 51 (history, key to species of North America); Granger, 1949: 18 (in key); Papp, 1960: 368, 377 (in key, key to species); Baltazar, 1962: 768 (in key); Mason, 1978: 722 (in key); Shenefelt, 1978: 1437 (catalogue, hosts); Marsh, 1979: 159 (catalogue of North America and Mexico); Marsh et al., 1987: 41 (in key); Quicke & Sharkey, 1989: 339, 345 (in key, notes, hosts); Wang et al., 2009: 32 View Cited Treatment (key to species of China); Li et al., 2020: 5 View Cited Treatment (key to species of China, diagnosis).

Type species. Ichneumon denigrator Linnaeus, 1758 .

Coelobracon Thomson, 1892: 1787 (name published in key (manipulus), type species assigned); Dalla Torre, 1898: 295 (synonymized). Type species: Ichneumon denigrator Linnaeus, 1758 .

Melanobracon Ashmead, 1900: 138 (new genus described in key, type species assigned); Szépligeti, 1904: 18 (synonymized). Type species: Bracon simplex Cresson, 1872 .

Atanycolidea Viereck, 1912: 617 (new genus, type species assigned); Shenefelt, 1943: 56 (synonymized). Type species: Bracon rugosiventris Ashmead, 1889 .

Diagnosis. Recognized within Atanycolus by the following combination of characters. Body medium–large sized. Scape highly modified and subcylindrical, base ventrally chamfer-shaped, apex ventrally expanded forward, bilaterally concave. Vein 1-SR+M of fore wing more or less straight, sometimes curved weakly. A single thickened basal bristle located at 1/5 of basal hind wing. First metasomal tergite divided into three parts, usually with different colors, the median part somewhat dark colored, second metasomal tergite with three smooth areas medio-basally and bilateral-basally. Ovipositor sheath longer or equal to body length. Hosts are woodborers of Buprestidae , Cerambycidae , and Curculionidae .

Key to Oriental and Palaearctic species similar to A. yangi sp. nov. in the genus Atanycolus

1 Female mesosoma totally black, male mesosoma black....................................................... 2

- Female mesosoma totally yellow to orange, male mesosoma most ( Fig. 22 View FIGURES 22–25 ) or totally black (see Ahmed et al. 2022: Fig. 1b View FIGURES 1–4 ) 6

2 Head red, only ocelli black (see Telenga 1936: key, redescription).......................... A. fulviceps (Kriechbaumer) View in CoL

- Head totally black, or yellow to red with frons or ocellar large field black......................................... 3

3 Head black, except for eye orbits yellowish red........................................... A. denigrator (Linnaeus)

- Head yellow to red, sometimes with stemmaticum and frons black widely or narrowly.............................. 4

4 Metasoma usually totally yellow, sometimes first metasomal tergite brown (see Li et al. 2020: Fig. 8e View FIGURES 5–9 ).................................................................................................... A. initiator (Fabricius)

- Metasoma with first tergite black (see Li et al. 2020: Figs 12j View FIGURES 10–12 , 18j View FIGURES 16–19 ); Triangular medio-basal area and antero-lateral areas of second tergite brown, other tergites yellow................................................................. 5

5 Head with stemmaticum and frons black, black area not extending backwards; temples weakly expanded behind eyes (see Li et al. 2020: key)..................................................................... A. setosus Li, He & Chen

- Head dorsally with a moderately large black and reverse guttiform spot around stemmaticum; temples strongly expanded behind eyes (see Li et al. 2020: key)....................................................... A. lindemani Tobias View in CoL

6 Female body length 13 mm, epipleuron of first metasomal tergite and sternum yellow, second metasomal tergite totally orange (see Ahmed et al. 2022: Figs 2g, 2h View FIGURES 1–4 )......................................... A. tangmargensis Ahmed, I. & Kazmi

- Female body length 7.5–10 mm, epipleuron of first metasomal tergite and sternum white, second metasomal tergite with marginal parts and sternum white, triangular medio-basal area and antero-lateral areas of second tergite black ( Figs 17 View FIGURES 16–19 , 24 View FIGURES 22–25 ).......................................................................... A. yangi Cao, Xie et Wang , sp. nov.