Abrodictyum pseudorigidum Bauret & Dubuisson (2016: 161)

Abrodictyum pseudorigidum Bauret & Dubuisson (2016: 161) View in CoL View Cited Treatment .

TYPE:— MADAGASCAR. Fianarantsoa: Ambatomboay, RNI d’Andringitra versant E, 40 km au Sud d’Ambalavao, sur les berges d’un affluent de la rivière Sahavatoy, 24 May 1995, F. Rakotondrainibe 2708 ( holotype P, P00059914!).

= Trichomanes rigidum Sw. var. lugubre H. Christ (1901: 849) View in CoL . Type:— DEMOCRATIC REPUBLIC OF CONGO. Kisantu, without date, J.B. Gillet s.n. ( holotype BR, BR 0000013468675!).

= Trichomanes rigidum Sw. var. annobonense Brause (1915: 378) View in CoL . Type:— EQUATORIAL GUINEA. Annobon Island: Summit of Santa Mina, 650 m, October 1911, G. W. J. Mildbraed 6700 ( holotype B, BW 20 0093175!).

Terrestrial ferns. Rhizomes short, erect, 0.5–2.0 cm in diameter, bearing long tufted erect red-brown trichomes, especially densely covering apex, and numerous robust roots. Fronds clustered, erect; stipes (2–)11–16(–26) cm long, cylindrical to mostly canaliculate, wingless or thinly winged on their upper part only, bearing numerous scattered trichomes identical to those present on rhizomes; rachis and main pinna axes with trichomes similar to those on the stipes and rhizomes. Laminae (8.0–)10.0–26.0(–31.0) × (3.0–)7.5–16.0(–19.0) cm, mostly triangular, less often sublanceolate to narrowly ovate, with more or less lanceolate to attenuated apex, tri-pinnate-pinnatifid, coriaceous and more or less opaque; pinnae (2.0–)3.5–7.0(–9.0) × 0.5–2.0(–3.5) cm, horizontal to oblique, basally sub-opposite, then alternate, lanceolate, or triangular to linear, and pinnate; pinnules pinnatifid to bipinnatifid (up to tripinnatifid for the largest specimens), lanceolate to linear; ultimate segments 0.2–0.4 mm wide, linear and single-veined with acute to rounded ends, the veins not reaching the margin; venation pinnate and anadromous. Sori paratactic, located mostly on acroscopic basal-most segments of proximal pinnules, 1.0–2.0 × 0.6–0.9 mm, 1–3(–6) per pinnule, up to 20 (sometimes more than 30) per pinna, free with a thin wing mostly at the base, cylindrical, truncated to slightly bilabiate with a slightly dilated mouth; receptacle short to long-exerted.

Distribution and habitat: —Sub-Saharan wet tropical Africa to western Indian Ocean (but absent from Mascarenes and Seychelles), terrestrial in the understory of rainforests from low to high elevations ( 200–2,500 m) ( Table 1).

Specimens examined: — COMOROS. Anjouan: 1875, M. Lavanchie s.n. (P00139638) ; May 1850, L.H. Boivin s.n. (P00139623); 1875, J.M. Hildebrandt 1793 (P00139637); Mjimandra , April 2011, E. Bidault 109 (P02439773) . Grande Comore: La Convalescence, versant ouest, November 2002, F. Rakotondrainibe 6771 (P00312031) ; Versant ouest du volcan Karthala, bois proche de Convalescence , October 2008, M. Pignal 3381 ( P02439785), M. Pignal 3391 ( P02439789) . Mohéli: Forêt de Saint Antoine , September 1995, N. Moulaert F.099 (P01485567) ; Chalet St Antoine , November 2008, M. Pignal 3473 (P00684891). Without locality: 1885, L. Humblot 1328 (P00139639, P01526320, P01526319, P00312496, P01430232, P01430231) ; without date, L.H. Boivin s.n. (P00301468).

— MADAGASCAR. Antsiranana: Befingotra, RS d’Anjanaharibe-Sud, sur le versant Sud-Est , October 1994, F. Rakotondrainibe 2097 (P0006321), 2097 bis (P0006322) ; November 1994, 2533 (P00046997); RNI 12 du Marojejy, au bord d’un affluent de la rivière Manantenina , October 1996, F. Rakotondrainibe 3374 (P00084841), 3390 (P00084866) ; Forêt de Betaolana , October 1999, F. Rakotondrainibe 4851 (P00179413) ; Forêt d’Analabe (massif d’Anjanaharibe-Sud ), October 1999, F. Rakotondrainibe 5052 (P00179699), 5092 (P00181052) ; PN de Marojejy, October 2001, F. Rakotondrainibe 6241 (P00243886). Fianarantsoa: Ambatomboay, RNI d’Andringitra versant Est , May 1995, F. Rakotondrainibe 2708 (P00059914) ; Réserve Spéciale d’Ivohibe , October 1997, F. Rakotondrainibe 4164 (P00134183), 4172 (P00134193), 4271 bis (P00134306) ; Corridor reliant les réserves d’Andringitra et d’Ivohibe , October 1997, F. Rakotondrainibe 4346 bis (P00134417) ; PN de Ranomafana, Forêt de Vatoharanana , October 2000, F. Rakotondrainibe 5916 (P00212397) ; Forêt de Vinanitelo , October 2000, F. Rakotondrainibe 6149 (P00212702) . Mahajanga: Daraina, Forêt de Binara , November 2001, F. Rakotondrainibe 6508 (P00248610) . Toamasina: Ambanizana, Piste menant au sommet d’Ambohitsitondroinan’ Ambanizana , sur le versant ouest, December 1993, F. Rakotondrainibe 1872 (P0006067) ; PN de Zahamena, Antanandava , January 2001, A. Rasolohery 241 (P00246411) ; PN de Zahamena, a 2 km d’Ankosy , le long de la SGR, June 2001, A. Rasolohery 502 (P00233447), 534 (P00233462) . Toliara: Eminiminy, Parcelle 1, R.N.I. n°11 d’Andohahela , October 1995, F. Rakotondrainibe 2893 (P00066991) ; Forêt d’Andranomay , December 1996, F. Rakotondrainibe 3791 (P00084658) ; Forêt d’Andrambovato , October 2000, F. Rakotondrainibe 5996 (P00212501), 5998 (P00212503) .