Knarr compressa ( Golemansky, 1979 ) Soler-Zamora & Useros & González-Miguéns & Gómez-Rodríguez & Lara, 2023

Soler-Zamora, Carmen, Useros, Fernando, González-Miguéns, Rubén, Gómez-Rodríguez, Pablo & Lara, Enrique, 2023, The problem of ‘ shadow species’ as illustrated with the taxonomic hotchpotch Cyphoderia ampulla (Rhizaria: Cyphoderiidae), Zoological Journal of the Linnean Society 199 (2), pp. 477-492 : 489-490

publication ID	https://doi.org/10.1093/zoolinnean/zlad040
persistent identifier	https://treatment.plazi.org/id/03C487FD-FFAC-FF9F-FC95-FD45FE7DFC9B
treatment provided by	Plazi
scientific name	Knarr compressa ( Golemansky, 1979 )
status	comb. nov.

Treatment

Knarr compressa ( Golemansky, 1979) comb. nov.

( Figs 1 View Figure 1 , 7 D View Figure 7 )

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Oleiformis Soler-Zamora, Useros, González-Miguéns & Lara gen. nov.

( Figs 1 View Figure 1 , 3 View Figure 3 , 5 View Figure 5 )

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Type species: Oleiformis major (Penard, 1890) comb. nov.

Description: Scales from the shell are non-imbricated, the species inhabits in freshwater ecosystems and they have a balloon-shaped shell that measures between 50–100 μm more than the largest species in the sister genus Cyphoderia .

Etymology: The genus name refers to the shape like an olive.

Included taxa: See Supporting Information, Appendix S1.

Oleiformis carmelae Soler-Zamora, Useros, González-Miguéns & Lara sp. nov.

( Figs 1 View Figure 1 , 5 B View Figure 5 2 View Figure 2 ; Model 2; Supporting Information, Fig. S1 View Figure 1 )

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Type locality: Morollón River , Fuente de la Gitana recreational area, La Peza, Spain ( 37º 51ʹ N 2º 47ʹ W) GoogleMaps .

Habitat: River sediment.

Measurements: 169.92–190.83 μm length, 67.44–83.14 μm width, 22.51–27.85 μm pseudostome large axis, and

2.65–2.99 × 1.93–2.26 μm scales from the middle of the shell ( N = 13).

Specific diagnosis: Elongated shell with an almost perfectly circular section. The shell is larger towards its middle part, giving an inflated appearance; rounded fundus. Short neck in comparison to other members of Cyphoderiidae . Rounded pseudostome with a thin organic layer surrounding it. Elliptical scales placed next to each other in a structured fashion; each scale is surrounded by six pores.

Intraspecific variability: Some individuals have a slightly more pointed fundus (Supporting Information, Fig. S1 View Figure 1 ).

Differential diagnosis: It can be confused with O. major because of their similar shell shape. O. carmelae is smaller, with relatively larger scales. It differs also genetically in both COI and SSU.

Etymology: In honour of Professor Carmen Zamora-Muñoz, mother of the first author.

Holotype: MA-Amoeba 11264.