Didymocarpus albicalyx C.B.Clarke

Didymocarpus albicalyx C.B.Clarke View in CoL in A.DC. & C.DC., Monogr. Phan. 5(1): 78. 1883. Lectotype (designated here): INDIA, East Himalaya , Sikkim,

s.d., Griffith 3839 ( K [ K00858157 digital image!], isolectotypes CAL [Acc. no 332191!], G [ G0041 8553, G00492372 digital images!], K [ K000939456 digital image!], M [ M0185736 digital image!], P [ P04060362 , P04060367 digital images!], S [ S110576 digital image!], W [ W0050224 digital image!]). Fig. 5 View Fig

Notes: Didymocarpus albicalyx was originally described by Clarke (1883) with the following diagnostic characters: “leaves ovate, serrate, and villous above; cymes many flowered, glabrous or sparsely pilose; calyx small, deeply lobed, white, with obovate, obtuse lobes; capsule 15 mm long, sub-sessile”. According to Clarke (1883, 1885 [“1884”]), the Sikkim Himalayan specimens described and illustrated as Didymocarpus villosus D.Don (as '' villosa '') by Clarke (1874) belonged to D. albicalyx , whereas D. villosus D. Don (1825) represented a distinct species presently known to occur only in Nepal and China.

In the protologue of D. albicalyx, Clarke (1883) cited three specimens: (i) Himalaya centralis, Sikkim, alt. 2000 m, Griffith 3839 (K, DC, G-BOISS); (ii) J.D. Hooker, s. n. (K, DC, P, W, G-BOISS, etc.), and (iii) Nepaul, prope flumen Tambur, J.D. Hooker, s. n. (K). In addition to these, Clarke (1883) cited '' Didymocarpus sp. n. 17 in Herbarium Indiae orientalis of J.D. Hooker & T. Thomson'', which also would qualify as the original materials of D. albicalyx . All the above specimens represent the syntypes of D. albicalyx (ICN Art. 9.6; Turland et al., 2018). The authors traced 25 sheets of these syntypes in nine herbaria (BM, CAL, G, K, M, MH, P, S and W). Amongst these, Griffith 3839 (K [K000858157]), with four fragments mounted on one sheet, the two upper left and right specimens with fruits, and the lower left and right ones with flowers and young fruits, was found a good match with Clarke’s original diagnosis and description of D. albicalyx . This sheet also carried Clarke’s annotation in his own handwriting: “ Didym . albicalyx C.B.Clarke, D. Don says inter alia that his villosa was large flowered and his type of villosa at the Mus. Brit. totally differs from this”. The sheet

Griffith 3839 in K is designated here as the lectotype and its duplicates in CAL, G, K, M, P, S and W as isolectotypes of D. albicalyx .