Alopoglossus angulatus, (LINNAEUS, 1758)

ALOPOGLOSSUS ANGULATUS ( LINNAEUS, 1758) View in CoL

[ FIGS 2–4 ( A. ANGULATUS ‘ EAST’); FIGS 5, 6, 7A;

TABLES 1 AND 2]

Lacerta angulata Linnaeus (1758: 204) View Cited Treatment .

Agama angulata Daudin (1802: 389) .

Alopoglossus carinicaudatus Ruthven (1924: 3 View in CoL , part); Ruibal (1952: 508, part); Cunha (1961: 123, part); Peters & Donoso-Barros (1970: 15, part).

Alopoglossus copii surinamensis Brongersma (1946: 231) View in CoL .

Alopoglossus angulatus Hoogmoed (1973: 216) View in CoL ; Ávila-Pires (1995: 307, part); Köhler et al. (2012: 176, part); Ribeiro-Júnior (2018: 29, his fig. 3B, 32, part).

Neotype: RMNH 15200 About RMNH , female, Suriname, Brown’s Mountain [Brownsberg] (4°56 ′ N, 55°10 ′ W), 20 August 1961, Marinus Hoogmoed GoogleMaps . Neotype selected by Hoogmoed (1973: 223), because the original holotype, from ‘America’, seems to be lost ( Figs 5, 6).

Material examined: Two hundred (N = 200) referred specimens for morphological analyses and 21 samples for molecular analyses from Brazil, French Guiana, Guyana and Suriname (see Supporting Information; Appendix S2, Table S1).

Diagnosis

Alopoglossus angulatus is distinguished from all other species of Alopoglossus by the combination of the following characters: (1) scales on sides of neck non-granular, keeled, imbricate (at least medial and posterior ones phylloid), in six to nine transverse rows; (2) three pairs of chin shields; (3) third pair of chin shields with rounded posterior margins, in direct contact with gulars or separated from them by a row of small scales; (4) absence of a pair of medial enlarged pregular scales; (5) scales on gular region subequal in size, but varying in shape: lateral ones strongly keeled, pointed, phylloid; anterior, medial and posterior ones varying from smooth to feebly keeled, phylloid; anteriormost transverse row with scales varying from smooth to feebly keeled, almost rounded posteriorly and varying from similar in size to slightly smaller than those in the proceeding rows; (6) ventral scales feebly keeled, mucronate, imbricate, with bluntly pointed posterior margins; (7) total number of femoral pores in males 20–29.