Phyllophaga (Cnemarachis) bretti Smith and Paulsen, 2015

Phyllophaga (Cnemarachis) bretti Smith and Paulsen View in CoL , new species

( Figs. 6–10 View Figs )

Type Series. Holotype, allotype, 156 paratypes. 92 males (40 CMNC [including holotype], 2 FCSA, 2 IESH, 2 MNHC, 39 UNSM, 7 USNM), 65 females (27 CMNC [including allotype], 2 FCSA, 2 IESH,

2 MNHC, 25 UNSM, 7 USNM) labeled: “ CUBA: Cienfuegos Prov. / Res. Ecológica Pico San Juan / 21.97083°N, 80.11859°W / 856 m; MV lights; forest edge / 18-V-2013; A.B.T.Smith ”. 1 female (UNSM) labeled: “ CUBA: Cienfuegos Prov. / east of Cuatro Vientos   GoogleMaps / 21.94596°N, 80.05318°W / 780 m; MV light; 19-V-2013 / A.B.T.Smith, F.Cala-Riquelme ”.

Diagnosis. Dorsal surface shiny, without pruinosity ( Fig. 6 View Figs ); clypeus sinuate; antennal club much shorter than funicle and scape; pronotal lateral margin entire; prosternum with obtuse projection basal to procoxae; metasternum setose; abdominal sternites shiny, without pruinosity, with some scattered setae; pygidium disc glabrous; male genitalia distinct as in Figs. 7–8 View Figs .

Holotype Description. Length 23 mm, width 11 mm. Dorsal surface shiny, without pruinosity; head brown, pronotum and elytra tan. Head: Surface densely punctate with deep, well-defined punctures; glabrous. Clypeus with apex reflexed, sinuate with middle distinctly shorter than sides; densely punctate. Frontoclypeal suture well-defined, clypeus and frons in same plane. Frons densely to moderately punctate. Antennae with 9 antennomeres; club slightly less than half length of funicle and scape; club, funicle, and scape shiny. Pronotum: Surface moderately punctate with deep, welldefined punctures; glabrous. Apical angles weakly acute; lateral margins entire (not crenulate), widest at medial point. Elytra: Surface moderately to densely punctate with shallow, well-defined punctures; appearing largely glabrous but with some scattered setae. Striae poorly defined, obscure. Legs: Protibia with 3 teeth, 2 nd tooth distinctly closer to apical tooth. Meso- and metatibiae without medial carinae, with spines on lateral margin. Metatibial spurs unmodified, shorter spur approximately 3/4 length of longer spur. Claws symmetrical with medial triangular tooth, apex unmodified. Venter: Prosternum with small, obtuse projection behind procoxae. Metasternum setose with long setae, without pruinosity. Abdominal sternites shiny without pruinosity, surface with scattered setae. Pygidium: Surface densely punctate with large, shallow, well-defined punctures; glabrous except with scattered setae apically and along lateral margins; shiny without pruinosity; color tan. Male genitalia: As in Figs. 7–8 View Figs .

Variation. Male length 21–24 mm, width 10–13 mm; female length 20–24 mm, width 9–12 mm. Terminal abdominal segment longer and more inflated in females than in males. Some males and most females with distinct, reddish brown longitudinal band medially on pronotum; when present, band half as wide as pronotum apically and tapering significantly towards base. Female genitalia are illustrated in Fig. 9 View Figs .

Etymology. This species is named in honor of Brett Ratcliffe (University of Nebraska State Museum, Lincoln, Nebraska, USA) in recognition of his support for this research project and the field expedition to Cuba. The patronym is derived from his first name to avoid confusion with Phyllophaga (Chlaenobia) ratcliffeiana Morón.

Distribution ( Fig. 10 View Figs ). CUBA: Cienfuegos: east of Cuatro Vientos, 21.94596°N, 80.05318°W (1); Reserva Ecológica Pico San Juan, 21.97083°N, 80.11859°W (157).

Temporal Data. May (158).

Remarks. This species keys to couplet 10 in Chapin (1932) but does not match the description or genitalic illustrations of Phyllophaga baracoana Chapin or any species resolved after this point in the key.