Phyllophaga (Cnemarachis) andersoni Smith and Paulsen, 2015

Phyllophaga (Cnemarachis) andersoni Smith and Paulsen View in CoL , new species

( Figs. 28–31 View Figs )

Type Series. Holotype, allotype, 83 paratypes. 32 males (13 CMNC [including holotype], 1 FSCA, 1 IESH, 1 MNHC, 15 UNSM, 1 USNM), 15 females (4 CMNC, 1 FSCA, 1 IESH, 1 MNHC, 8 UNSM) labeled: “ CUBA: Cienfuegos Prov. / Res. Ecológica Pico San Juan   GoogleMaps / 21.97083°N, 80.11859°W / 856 m; MV lights; forest edge / 18-V-2013; A.B.T.Smith ”. 14 males (7 CMNC, 5 UNSM, 2 USNM), 10 females (5 CMNC [including allotype], 3 UNSM, 2 USNM) labeled: “ CUBA: Cienfuegos Prov. / Soledad   GoogleMaps , 71m / 22.12682°N, 80.33289°W / 21-V-2013; MV light / R.Anderson, G.Zhang ”. 13 males (CMNC) labeled: “ CUBA: Cienfuegos / Mayari , 1.5 km E. / 21.97100 -

80.11644, 866m / 18.v.2013, R.Anderson / 2013- 020X, hand collections”. 1 male (CMNC) labeled: “ CUBA: Cienfuegos / Mayari   GoogleMaps , 1.5 km E. / 21.97100 - 80.11644, 866m / 18.v.2013, R.Anderson / 2013-020, hardwood forest litter”.

Diagnosis. Elytra pruinose (most males) ( Fig. 28 View Figs ) or shiny without pruinosity (some males and all females) ( Fig. 29 View Figs ); clypeus sinuate; antennal club much shorter than funicle and scape; pronotal lateral margin entire; prosternum with two spines basal to procoxae; metasternum glabrous with a few scattered setae adjacent to midline; abdominal sternites opaque/pruinose laterally, glabrous medially, sternite 5 with much wider pruinose patch; pygidium disc glabrous; male genitalia distinct as in Fig. 30 View Figs .

Holotype Description. Length 14 mm, width 7 mm. Dorsal surface shiny, dark brown with purplish gray, pruinose reflections on elytra (small patches at humeral and apical umbones of elytra without pruinosity) ( Fig. 28 View Figs ). Head and pronotum shiny, without pruinosity. Head: Surface densely to moderately punctate with deep, well-defined punctures; glabrous. Clypeus with apex reflexed, sinuate with middle distinctly shorter than sides; densely punctate. Frontoclypeal suture well-defined, clypeus and frons in same plane. Frons moderately punctate. Antennae with 9 antennomeres; club approximately 1/3 length of funicle and scape; club, funicle, and scape shiny. Pronotum: Surface moderately punctate with deep, well-defined punctures; glabrous. Apical angles moderately acute; lateral margins entire (not crenulate), widest slightly apical to medial point. Elytra: Surface moderately to densely punctate with deep, well-defined punctures; largely glabrous but with some scattered setae especially towards apex. Striae poorly defined, obscure. Legs: Protibia with 3 teeth, 2 nd tooth distinctly closer to apical tooth. Meso- and metatibia without medial carinae, with spines on lateral margin. Metatibial spurs unmodified, shorter spur approximately 3/4 length of longer spur. Claws symmetrical with medial triangular tooth, apex unmodified. Venter: Prosternum with 2 small spines behind procoxae. Metasternum glabrous (except for a few scattered setae adjacent to midline), medially shiny and laterally with opaque/pruinose coating. Abdominal sternites medially shiny and laterally with opaque/pruinose coating (pruinosity much wider on sternite 5). Pygidium: Surface moderately to densely punctate with large, well-defined punctures; glabrous; shiny with pruinosity along basal margin; color brown medially and light brown around all margins. Male genitalia: As in Fig. 30 View Figs .

Variation. Male length 11–14 mm, width 6–8 mm; female length 12–13 mm, width 7–8 mm. Terminal abdominal segment longer and more inflated in females than in males. Two males and all females had no elytral pruinosity ( Fig. 29 View Figs ). Most males had pruinose elytra as in holotype.

Etymology. This species is named in honor of Robert Anderson (Canadian Museum of Nature, Ottawa, Ontario, Canada) in recognition of his support for this field expedition and his research on the weevils of Cuba.

Distribution ( Fig. 31 View Figs ). CUBA: Cienfuegos: Mayari (1.5 km E), 21.97100°N, 80.11644°W (14); Reserva Ecológica Pico San Juan, 21.97083°N, 80.11859 ° W (47); Soledad, 22.12682 ° N, 80.33289°W (24).

Temporal Data. May (85).

Remarks. This species does not effectively key past couplet 21 in Chapin (1932) and does not match the description or genitalic illustrations of any species resolved after this point in the key.