Puto, Signoret

Key to adult females of Puto View in CoL View at ENA from the Old World (updated from Danzig & Gavrilov 2014)

Notes: The key does not include P. brunnitarsis ( Signoret 1875) because the available description is inadequate. The data of P. barberi (Cockerell) , P. marsicanus and P. thailandicus are derived from Williams & Granara de Willink (1992), Marotta & Tranfaglia (1993) and Williams (2004) respectively.

1(0) Claw with denticle well developed........................................................................ 2

- Claw without denticle............................................................... P. borealis (Borchsenius) View in CoL

2(1) Dorsal tubular ducts present............................................................................. 9

- Dorsal tubular ducts absent............................................................................. 3

3(2) Ventral tubular ducts of 2 or 3 sizes....................................................................... 4

- Ventral tubular ducts all same size........................................................................ 6

4(3) Ventral tubular ducts of 2 sizes........................................................................... 5

- Ventral tubular ducts of 3 sizes............................................................ P. orientalis Danzig View in CoL

5(4) Large ducts present along margins of thorax and posterior abdominal sternites, absent from median areas of posterior abdominal sternites........................................................................... P. antennatus (Signoret) View in CoL

- Large ducts absent from margins of thorax and posterior abdominal sternites, but present on median areas of posterior abdominal sternites............................................................................. P. barberi (Cockerell) View in CoL

6(3) Ventral tubular ducts scattered throughout posterior part of body, but absent from anterior part of body... P. konoi Takahashi View in CoL

- Ventral tubular ducts present in frontal group only, or (rarely) totally absent....................................... 7

7(6) Multilocular disc-pores numerous, scattered on all medial zone of venter.................. P. superbus (Leonardi) View in CoL , partly

- Multilocular disc-pores few, present on cephalothoracic sternites only or totally absent.............................. 8

8(7) Multilocular disc-pores present on cephalothoracic sternites............................... P. peyerimhoffi (Vayssiere) View in CoL

- Multilocular disc-pores absent.......................................................... P. israelensis Ben-Dov View in CoL

9(2) Dorsal tubular ducts of 2 sizes; smaller ducts present along margins of abdominal tergites.......... P. caucasicus Hadzibejli View in CoL

- Dorsal tubular ducts all same size, large.................................................................. 10

10(9) Dorsal tubular ducts wide; diameter of their openings more than 2 times larger than diameter of a trilocular pore......... 11

- Dorsal tubular ducts narrow; diameter of their openings about the same size as diameter of a trilocular pore............. 13

11(10) Dorsal tubular ducts located only in cerarii. Ventral tubular ducts all same size, present only in frontal group................................................................................................ subericola (Vayssiere)

- Dorsal tubular ducts located between cerarii. Ventral tubular ducts of 2 or 3 sizes, numerous on all abdominal sternites.... 12

12(11) Quinquelocular disc-pores present....................................................... P. huangshanensis Wu

- Quinquelocular disc-pores absent........................................................... P. tubulifer Danzig View in CoL

13(10) Dorsal tubular ducts present along body margin only........................................................ 14

- Dorsal tubular ducts present in medial zones of tergites as well as along margins.................................. 15

14(13) Dorsum with additional cerarii, forming 6 longitudinal rows. Ventral margins without small oral collar tubular ducts.................................................................................... P. superbus (Leonardi) View in CoL , partly

- Dorsum without additional cerarii. Ventral margins with groups of small oral collar tubular ducts.................................................................................................. P. sinensis Zheng & Wu , sp. n.

15(13) Ventral tubular ducts present on thoracic and abdominal sternites as well as in frontal group......................... 16

- Ventral tubular ducts absent from thoracic and abdominal sternites, present only in frontal group..................... 17

16(15) Dorsum with additional cerarii forming 2 longitudinal rows, plus several additional cerarii on abdominal segment VI..................................................................................... P. megriensis (Borchsenius) View in CoL

- Dorsum without additional cerarii...................................................... P. thailandicus Williams

17(15) Dorsum with additional cerarii forming 6 longitudinal rows........................ P. marsicanus Marotta & Tranfaglia View in CoL

- Dorsum with additional cerarii forming 2 longitudinal rows.............................. P. kondarensis (Borchsenius) View in CoL