Horizon

Horizon and age. – Upper Cretaceous ~98.79 ± 0.62 million years old ( Shi et al., 2012).

Discussion. – This species was recently described and taxonomically associated with Burmacypha longicornis to form the subfamily Iotacyphinae Shcherbakov et al., 2024 . This clade can easily be identified by several morphological features. First, the forewings have intricate, small tessellating hexagonal patterns. Additionally, the forewings have an apical fusion of MP with CuA. This unique character creates an enclosed, large central cell on the forewings, a feature not known from any other †Lophioneurida.

Currently the genus is monotypic. See the generic differentiation below for differentiation from the other two genera presently known in the clade; Burmacypha longicornis and Retiptera brennae gen. et sp. nov. Additional specimens of Iotacypha zherikhini reviewed herein ( Fig. 1, 2) are very similar to the type material from the original description. Due to the taphonomy of these additional specimens, no additional morphological details can be added to the species description.