Davidina, OBERTHUR, 1879

GENUS DAVIDINA OBERTHÜR, 1879 View in CoL

Et. Entom. 4: 19, t. 2, fig. 1. Type-species: Davidina armandi Oberthür, 1879 (Et. Entom. 4: 19, t. 2, f. 1).

= Leechia Röber, 1907 . In Seitz, Grossschmetterlinge der Erde 1: 43, t. 19b. Type-species: Leechia alticola Röber, 1907 (In Seitz, Grossschmetterlinge der Erde 1:43, t. 19b).

= Sinosatyrus Lee Chuan-Lung, 1988. Chinese Science Bull. 33 (4): 320. Type-species: Sinosatyrus beyingensis Lee, 1988.

= Protoeneis P. Gorbunov, 2001. In Gorbunov, The butterflies of Russia: classification, genitalia, keys for identification ( Lepidoptera : Hesperioidea and Papilionoidea): 228. Type-species: Chionobas nanna Ménétriés, 1858 (Bull. Class. phys.-math.Acad. imp. Sci. St. Petersb. 17(12/14): 216).

1. Davidina armandi Oberthür, 1879 . Et. Entom. 4: 19, t. 2, fig. 1. TL: Pe-hoa-tschan, China. Holotype in BMNH ( Fig. 1 View Figure 1 ).

= alticola (Röber, 1907) ( Leechia alticola ). In Seitz, Grossschmetterlinge der Erde 1: 43, t. 19b. TL: ‘Central-China (Tschang-yang)’. TM in BMNH.

= beyingensis (Lee Chuan-Lung, 1988) (Sinosatyrus beyingensis). Chinese Science Bull. 33 (4): 320.

Distribution: N. E. China.

Note. Lee (1988) spelled his new species of Sinosatyrus in two different ways: beÿingensis and beÿiengensis (both on p. 320, note the diaeresis mark placed over the ‘y’). Both spellings appear to be misprints of the letters ‘ ij ’, thus possibly Lee’s original intention was to name the species beijingensis but that was not what was printed. Here, acting as First Reviser, we select beyingensis as the correct original spelling, simultaneously deleting the diacritic mark, following Arts. 32.2.1 and 32.5.2.1 of the ICZN.

2. Davidina diluta (Lukhtanov, 1994) ( Oeneis diluta ). In Lukhtanov & Lukhtanov, Die Tagfalter Nordwestasiens, Herbipoliana 3: 138, t. 25, fig. 6. TL: Susch, Ujukski-Gebirge, Tuva, S. Siberia, Russia. HT in ZISP .

Distribution: Russia: C. Tuva.

3. Davidina lederi (Alpheraky, 1897) ( Oeneis Tarpeia Pall. var. Lederi ). Mém. Lép. Rom. 9: 196. TL: ‘ Urga’ [Ulanbator], Mongolia (probably in the Khangai Mts). LT in ZISP, designated by U. Eitschberger & V.A.Lukhtanov (1994), Atalanta 25:163 .

= sapozhnikovi Korshunov, 1982 ( Oeneis sapozhnikovi). Gel’minty, kleshchi i nasekomye: 86, figs 1–2. TL – 10 km NW Mungen-Mort, Central Aimak, Mongolia. HT in SZM.

Distribution: N. and C. Mongolia. Russia: Irkut River valley in S. Siberia, S. Tuva.

4. Davidina mongolica (Oberthür, 1876)

4a. Davidina mongolica mongolica (Oberthür, 1876) ( Chionobas mongolica ). Et. Entom. 2: 31, t.4, fig. 6. TL: ‘ Mongolie orientale. Elle vole en été dans les montagnes, à une altitude moyenne de 500 mètres’ TM in BMNH.

= tsingtaua Austaut, 1911 ( Oeneis mongolica tsingtaua ). Ent. Zeitschr. 24: 244. TL: Tsingtau, Shantung, China or. TM -?.

= tsingtauana auct. (misspelling).

= mandschurica O.Bang-Haas, 1939 ( Oeneis mongolica mandschurica ). Iris 53: 49, t.1, fig. 6. TL: Kintschou, Prov. Fengtien, Manchuria mer. or., China. TM in ZMHU.

f. extincta O. Bang-Haas, 1939 . Iris 53:49. TL: Kintschou, Prov. Fengtien, Manchuria mer. or., China. TM in ZMHU.

Distribution: N. E. China: Inner Mongolia, Liaoning.

4b. Davidina mongolica hoenei (Gross, 1970) ( Oeneis mongolica hoenei ). Mitt. Münch. Ent. Ges. 58: 22. TL: ‘Mienshan, Shansi, Nordchina’. HT in MAKB.

Distribution: N. China: Shansi [Shanxi].

4c. Davidina mongolica coreana (Matsumura, 1927) ( Oeneis nanna coreana ). Ins. Mats. 1: 163, t. 5, fig. 9. TL: ‘Genzan’, Korea. TM in HUS (?).

= okamotonis Matsumura, 1927 (‘ Oeneis nanna coreana okamotonis n. ab.’, p. 164; ‘ Oeneis nanna okamotonis ’, p. 160). Ins. Mats. 1: 164, t. 5, f.3. TL: ‘Suigun’, Korea. TM in HUS (?).

f. fumosa O. Bang-Haas, 1939 . Iris 53:49, t.1, f.6. TL: ‘ Corea int., bei Gensan’. TM in ZMHU.

Distribution: N. Korea.

4d. Davidina mongolica walkyria (Fixsen, 1887) ( Oeneis walkyria ). Mém. Lép. Rom. 3: 310. t.14, fig. 4. TL: ‘in der Umgebung Pung-Tungs’, Korea. TM in ZISP.

= shonis Matsumura, 1927 (‘ Oeneis nanna walkyria shonis n. ab.’, p. 163; ‘ Oeneis nanna shonis ’, p. 150). Ins. Mats. 1: 163, t. 5, fig. 2. TL: ‘Mt. Daitoku’, Korea. TM in HUS (?).

f. hakuba Doi, 1934 ( Oeneis nanna walkyria f. hakuba). In Mori, Doi & Cho, Coloured butterflies from Korea (2): 16, t. 12, fig. 4. TL: Korea. TM -?.

f. soibona Doi, 1934 ( Oeneis nanna walkyria f. soibona). In Mori, Doi & Cho, Coloured butterflies from Korea (2): 16, t. 12, fig. 7. TL: Korea. TM -?.

= ? masuiana Matsumura, 1919 View in CoL . Thous. Ins., Add. 3: 547, t. 38, f. 2. Ins. Mats. 1:160 (1927). TM-HUS (?).

Distribution: C. Korea.

4 e. Davidina mongolica hallasanensis (Murayama, 1991) ( Oeneis urda hallasanensis ). Nature and Insects 26 (3): 20. TL: Mt. Hallasan, Cheju-do Island, S. Korea. TM -?.

Distribution: Korea: Cheju-do Island.

5. Davidina nanna (Ménétriés, 1858)

5a. Davidina nanna dzhulukuli (Korshunov, 1998) ( Oeneis anna dzhulukuli ). In Korshunov , Novye opisaniya i dopolneniya dlya knigi ‘ Dnevnye babochki Aziatskoi chasti Rossii’ : 27, t. 17(4). TL: Taboshak Mt., Kuraiski Chrebet, Altai, Russia. TM in SZM .

Distribution: Russia: S. E. Altai (Kurai, S. Tchuja and Sailjugem Mountains, Ukok-Plateau), W. Tuva.

5b. Davidina nanna anna (Austaut, 1911) ( Oeneis nanna var. anna ). Ent. Zeitschr. 24: 243. TL: ‘Arasagun-Gol’ (N. Mongolia). TM in?.

= brunhilda A. Bang-Haas, 1912 ( Oeneis brunhilda ). Iris 26:105, t. 6, f. 1. TL: ‘aus dem Sajan-Gebiete’. Acording to the syntype labels the type-locality is ‘ Sajan, Arasagun-Gol’ (N. Mongolia) and ‘ Sajan , Arsyn’ ( Russia, S. Tuva). STs in ZMHU.

= burjatica Korshunov et Nikolaev, 2002 ( Oeneis anna burjatica ). In Korshunov, 2002, Bulavousye Cheshuekrylaye Severnoi Azii (Butts N Asia): 319.

D i s t r i b u t i o n: R u s s i a: S. T u v a, S. B u r y a t i a. N. Mongolia.

5c. Davidina nanna nanna (Ménétriés, 1858) ( Chionobas nanna ). In Bull. Class. phys.-math.Acad. imp. Sci. St. Petersb. 17(12/14): 216. TL: ‘Amour’ [Amur], Russia. HT in ZISP .

= hulda Staudinger, 1887 ( Oeneis hulda ). Mém. Lép. Rom. 3: 149, t.16, f.8. TL: Pokroffka , Amur, Russia. LT in ZMHU, designated by F. J. Gross (1970), Mitt. Münch. Ent. Ges. 58: 20, Abb. 29.

= coriacea Seitz, 1909 ( Oeneis nanna coriacea ). In Seitz, Grossschmetterlinge der Erde 1:121, t. 40g. TL: ‘Apfelgebirge’, S. Siberia, Russia. TM in SMF.

Distribution: Russia: Transbaikalia, Amur basin. N. E. Mongolia. N.E. China.

5d. Davidina nanna jakutski (Korshunov, 1998) ( Oeneis jakutski ). Novye opisaniya i dopolneniya dlya knigi ‘Dnevnye babochki Aziatskoi chasti Rossii’: 27, t. 17(3). TL: Yakutsk, botsad [botanical garden], Yakutia, Russia. TM in SZM.

Distribution: Russia: Yakutia.

5e. Davidina nanna dzhugdzhuri (Sheljuzhko, 1929) ( Oeneis dzhugdzhuri ). Mitt. Münch. Ent. Ges. 19: 349, t. 28, figs. 3–4. TL: ‘in der Bergkette Dzgugdzhur (Grenze der Provinzen Amur und Jakutsk), an den Quellen des Flusses Dzhelinda’ [Yakutia], Russia. HT in coll. Sheljuzhko (Kiev University, Ukraine).

Distribution: Russia: Southern and Eastern Yakutia, northern part of Amurskaya oblast’.

5f. Davidina nanna taimyrica (Lukhtanov & Eitschberger, 2001) . ( Oeneis nanna taimyrica ). In: Lukhtanov & Eitschberger (2001) Catalogue of the genera Oeneis and Davidina (Bauer E, Frankenbach T, eds) Butterflies of the World, Supplement 4: 33.

TL: Russia, Taimyr Peninsula, Putorana plateau, Ayan Lake, near Ayan River. HT in Zoological Museum of Moscow University.

Distribution: Russia: Northern Siberia (Taimyr Peninsula).

6. Davidina sculda (Eversmann, 1851)

6a. Davidina sculda sculda (Eversmann, 1851) [ Hipparchia (Chionobas) sculda ]. Bull. Soc. Nat. Mosc. 24: 612. TL: ‘environs de Kiachta de la Sibérie orientale’ (Buryatia, S. Siberia, Russia). STs in ZISP .

= sculda Keferstein, 1851 , Stett. ent. Ztg. 12(9): 274 (a nomen nudum) (Chionabas [sic])

= velleda Austaut, 1911 ( Oeneis velleda ). Int. Ent. Zeitschr. 5: 360. TL: Siberia. TM in?.

Distribution: Russia: Altai, Tuva, Sayan Mts,

S. Buryatia. N. E. Mongolia.

6b. Davidina sculda pseudosculda (Korshunov, 1977) ( Oeneis pseudosculda ). Insects of Mongolia 5: 662, fig, 1. TL: Sudzykte, Central Aimak, Mongolia. HT in ZISP.

Distribution: C. Mongolia.

6c. Davidina sculda pumila (Staudinger, 1892) ( Oeneis Sculda Ev. var. Pumila Stgr. ). Mém. Lép. Rom. 6: 201. TL: ‘Pokr.[offka]’ and ‘am oberen Amur’ (Transbaikalia region, Russia). STs in ZMHU.

Distribution: Russia: Transbaikal (Tchita region), Amur region. N. E. Mongolia. N. E. China.

6d. Davidina sculda vadimi (Korshunov, 1995) ( Oeneis sculda vadimi ). In Korshunov & Gorbunov, Dnevnye babochki Asiatskoi chasti Rossii: 138. TL: Severobaikalsk, Verkhneangarski Khrebet, N. Transbaikal, Russia. HT in SZM.

Distribution: Russia: N. Transbaikal, Yakutia.

7. Davidina tarpeia (Pallas, 1771)

7a. Davidina tarpeia tarpeia (Pallas, 1771) ( Papilio tarpeia ). Reise versch. Prov. russ. Reiches 1: 171 (tarpeja), 480 ( tarpeia ). TL: ‘In campis aridis ad Volgam copio fus Maio’, Volga region, Russia. TM lost.

= celimene (Cramer, 1782) ( Papilio celimene ). Pap. Exot. 4: 169, t. 375 E, F. TL: ‘Siberie’.

= vacuna Grum-Grshimailo, [1891] ( Oeneis Vacuna ). Horae Soc. Ent. Ross. 25: 458. TL: ‘in montibus ad Dongar-tschen’, S. of Xining , Qinghai, China. HT in BMNH.

= rozhdestvenskyi Korb et Yakovlev , 1997 ( Oeneis tarpeia rozhdestvenskyi ). Zoosystematica Rossica 5 (2):313. TL: Shtabka, Altai, Russia. HT in ZISP .

= ukokana Korb, 1998 ( Oeneis tarpeia ukokana ) Alexanor 20: 144, figs 1–3. TL: ‘Südaltai, Ukok, Dschasator’, Russia. HT in ZISP.

= baueri Lukhtanov & Eitschberger, 2000 ( Oeneis tarpeia baueri ). Schmetterlinge der Erde 11: 8. TL: Mezen’ r.[iver], vic. Sush’, Tuva, Siberia, Russia, 1100– 1200 m. HT in McGuire Center for Lepidoptera and Biodiversity, University of Florida.

Distribution: Russia: E.and S. E. Europe, N. Caucasus (Kislovodsk, Teberda, Elbrus), S. Ural, S. W. Siberia (W. Siberian Lowland), Altai, Tuva. N. and E. Kazakhstan. W. Mongolia. NW. China.

Note. There are two different spellings of this name in the original description and in subsequent works. Here we select tarpeia as the most common spelling in historical and contemporary works.

7b. Davidina tarpeia grossi (Eitschberger & Lukhtanov, 1994) ( Oeneis tarpeia grossi ). Atalanta 25: 164, t. Va, figs 1–3. TL: ‘ 20 km NW Selenduma’ , Buryatia, S. Siberia, Russia. HT in ZISP .

Distribution: Russia: Buryatia, Chita region. N. Mongolia.

8. Davidina uhleri (Reakirt, 1866)

8a. Davidina uhleri varuna (W. H. Edwards, 1882) ( Chionobas varuna ). Canadian Ent. 14: 205. TL: ‘plains of Dacotah Terr.’, Canada. LT in CM.

ab. dennisi Gander, 1927 . Canadian Ent. 59: 285– 286 (1927). TL: Beulah, Manitoba. HT in CNC.

Distribution: Canada: British Columbia, Alberta, Saskatchewan, Manitoba. USA: Montana, Dakotas , Nebraska, W. Minnesota.

8b. Davidina uhleri nahanni (Dyar, 1904) ( Oeneis nahanni ). Proc. Ent. Soc. Washington 6:142. TL: Nahanni Mts., Dist. of Mackenzie, Canada. HT in USNM.

Distribution: Canada: North-West Territories (Mackenzie Mts).

8c. Davidina uhleri cairnesi (Gibson, 1920) ( Oeneis cairnesi ). Rept. Canadian Arctic Exped. 3 (1): 15. TL: White River dist. , Yukon Territory, Canada. HT in CNC .

= kluanensis (Hassler, 2002) ( Oeneis nanna kluanensis ). Nachricht. Entomol. Ver. Apollo 22: 197– 205. TL: SW-Yukon, NW Burwash Landing, Quill Creek Area. HT in SMF.

Distribution: Canada: Yukon, northwestern corner of the North-West Territories. USA: N. E. Alaska.

8d. Davidina uhleri uhleri (Reakirt, 1866) ( Chionobas Uhleri ). Proc. Ent. Soc. Philadelphia 6:143. TL: ‘ Rocky Mts. , Colorado Territory’ , USA, restricted to vic. Georgetown, Clear Creek Co., Colorado by F. M. Brown (1953), American Mus. Novitates (1625): 6. TM in FMNH .

= obscura (W. H. Edwards, 1892) ( Chionobas Uhleri var. Obscura ). Butt. N. America 3:294. TL: Clear Creek Canyon, Jefferson Co., Colorado. HT in CM.

Distribution: USA: Colorado east of the Continental Divide.

8e. Davidina uhleri reinthali (F. M. Brown, 1953) ( Oeneis uhleri reinthali ). American Mus. Novitates (1625): 10–11. TL: Gothic, Colorado. HT in AMNH.

Distribution: USA: western slope of the Continental Divide in Colorado.

9. Davidina urda (Eversmann, 1847)

9a. Davidina urda urda (Eversmann, 1847) ( Hipparchia urda ). Bull. Soc. Nat. Mosc. 20 (3): 69, t. 2, f. 1–4.TL: ‘Dauria’, Russia. TM in ZISP.

= umbra Staudinger, 1892 ( Oen. urda Ev. var. Umbra ). Iris 5: 335. TL: ‘Südamur’, Russia. TM in ZMHU.

= laeta Austaut, 1908 ( Oeneis urda Evers. var. laeta ). Ent. Zeitschr. 22: 44. TL: ‘Sayan’. TM -?.

= trybomi Bryk, 1946 ( Oeneis urda tribomi ). Arkiv Zool. 38A (3): 25, t. 4, fig. C3. TL: ‘Krasnojarsk’, [S. Siberia, Russia]. HT in NRS.

= rasa Bryk, 1946, Arkiv Zool. 38A (3): 24 (an infrasubspecific name).

ab. albidior Austaut, 1908 . Ent. Zeitschr. 22: 44. TL: ‘Sayan’. TM -?

ab. banghaasi Austaut, 1908 . Ent. Zeitschr. 22: 44. TL: ‘Sayan’. TM -?

Distribution: Russia: N. Altai (Tongosh Mountains), S. Siberia, Transbaikal, Amur and Primorye regions, S. and Central Yakutia. N. Mongolia. N. E. China.

9b. Davidina urda tschiliensis (O. Bang-Haas, 1933) ( Oeneis urda tschiliensis ). Ent. Zeitschr. 47: 98. TL: Hsingan mont., Tunkia-jingze, Prov. Tschili, China sept. HT is probably in ZMHU.

Distribution: N. E. China: Hebei.

9c. Davidina urda monteviri Bryk, 1946 ( Oeneis urda monteviri ). Arkiv Zool. 38A (3): 24, t. 4, fig. A3. TL: ‘Shinten’, Korea. HT in NRS(?).

Distribution: N. Korea.