Hymenocardia Wall. ex Lindl.

Hymenocardia Wall. ex Lindl. View in CoL

Hymenocardia Wall. ex Lindl. (1836) View in CoL 441; Endl. (1837) 288 (nr. 1899); Tul. (1851) 256; Baill. (1857) 994; (1858) 599,pl. 27:f. 24, 25;Müll.Arg. (1866) 476; Kurz (1877) 394; Benth. (1880) 285; Hook.f. (1887) 376; Pax (1890) 30;Pax & K.Hoffm. (1922) 72;Gagnep.(1927) 543;Pax & K.Hoffm. (1931) 51; Airy Shaw (1964) 262; (1972a) 363; Whitmore (1973) 103; Radcl.-Sm. (1987) 52; G.L.Webster (1994) 53; Govaerts et al. (2000) 1013; Radcl.- Sm. (2001) 74; Welzen (2007) 341; G.L.Webster (2014) 82,f. 15. ― Type: Hymenocardia punctata Wall. ex Lindl. View in CoL

Samaropyxis Miq. (1860) View in CoL 464. ― Type: Samaropyxis elliptica Miq. View in CoL (= Hymenocardia punctata Wall. ex Lindl. View in CoL ).

Shrubs to trees, dioecious, deciduous, leaves appearing during flowering; branches hairy with scale-like hairs and simple hairs when young. Indumentum of simple and scale-like hairs (re- corded as glandular). Stipules triangular, thick, early caducous. Leaves distichous (to opposite) on branches, simple; petiole not pulvinate, reniform in transverse section; blades elliptic, margin entire, glabrous above to hairy on midrib with simple hairs, hairy beneath, venation densely reticulate, few nerves, latter looped and closed near margin. Inflorescences dense spikes

276 Blumea – Volume 61 / 3, 2016

1 mm e d 1 mm i c 1 mm j 1 mm g b 1 mm h f 1 mm 1 cm a

to panicles, axillary and catkin-like when staminate, terminal and few-flowered racemes when pistillate; flowers bracteate. Flowers actinomorphic; petals and disc absent. Staminate flowers: pedicel very short; calyx mainly 5-lobed, cupular, lobes triangular, imbricate to almost valvate; stamens mainly 5, episepalous, basally united or free, anthers very large, 2-thecate, dorsifixed, horizontal when dehiscing extrorse via lengthwise slits, connective with subapical, dorsal gland; pistillode on top of androphore, a short cylinder, apically slightly 2-lobed. Pistillate flowers shortly pedicelled, sepals (4–)5(–8), free, longtriangular, valvate, soon caducous leaving cup-shaped scars; ovary 2-locular, flattened; ovules 2 per locule; style absent or minute, stigmas 2, unlobed, long papillate. Fruits flat, 2-lobed, usually heart-shaped, samara-like capsules (rhegmas), winged or not. Seeds usually 1 per locule developed, naked.

Distribution ― A genus of six species ( Govaerts et al. 2000), of which five in continental Africa and one in Southeast Asia main land and W Malesia.