Cenomanisega, Lucena & Melo, 2025

Genus † Cenomanisega gen. nov.

ZooBank registration: LSID: urn:lsid:zoobank.org:act:

Type species: † Cenomanisega krombeini sp. nov.

Diagnosis

Male mandible elongated, flattened, broadly foliaceous, lacking subapical tooth, apparently truncate apically; female mandible delicate, tapering apically, without subapical teeth on laminar edge; general body shape long and flattened dorsoventrally; pronotum long, slightly longer than scutum in mid-line, posterolateral lobe well separated from tegula, lacking posterolateral and posteromedial pits; mesopleuron anteriorly delimited with carina, mostly evenly convex, with broad posterior depression below scrobal pit, without transverse sulcus; posterolateral corner of propodeum round, lacking posterior propodeal projection, posterior surface declivitous.

Description

Head: Pyriform, front well developed and protruding; eye bulging, occupying much of lateral head, following contour of frons anteriorly; lower gena 0.4 × eye width, upper gena narrowed, nearly indistinct; malar space slightly shorter than 0.4 × eye height, crossed by longitudinal malar sulcus; pre-hypostomal foveae absent; occipital carina extended dorsally, fading gradually laterally; scapal basin shallowly excavated, disc smooth, borders smoothly round, without carina or sharp edges; scape length 4.1 × maximum width; labial and maxillary palpomeres with three and four articles, respectively; flagellum fusiform, with intermediate articles broader than long, progressively shortened from F4 to F10 and with flattened ventral areas on F4–F11; mandible strongly tapering apically, main axis straight, lacking subapical teeth on laminar edge.

Mesosoma: Elongated and somewhat fusiform, slightly flattened dorsally; pronotum long, slightly longer than scutum in mid-line, posterior border concave, crossed anteriorly and posteriorly by transverse grooves, posterolateral lobe well separated from tegula, lacking posterolateral and posteromedial pits; notaulus deeply impressed, strongly converging posteriorly; parapsidal line well marked posteriorly; scutellum slightly shorter than half length of scutum in mid-line, simple, evenly convex, without anterior transverse groove, pits, or foveae; mesopleuron anteriorly bordered with carina, evenly convex, with wide posterior shallow depression below scrobal pit, without transverse sulcus; metanotum about one-third of scutellum in mid-line, expanded and slightly depressed laterally, without differentiated disc; propodeum box-like, dorsum longer than high, with distinguishable lateral, dorsal, and posterior surfaces; dorsal and posterior surfaces medially crossed by contiguous longitudinal groove; dorsum and posterior surface delimited laterally by confluent grooves, metapostnotum indicated anteriorly as a distinct transverse raised band, posterolateral corner round, lacking posterior propodeal projection, posterior surface declivitous; tarsal claw with short, perpendicular, submedial tooth; inner metatibial spur distinctly shorter than the outer; dorsum of metacoxa with strongly produced longitudinal carina; dorsum of metafemur evenly convex, without anterior expansion; costal cell narrow, first abscissa of M diverging before 1cu-a, gently and smoothly curved submedially; Rs+M arising apically on M; medial cell indicated by spectral and nebulous veins; R1 slightly shorter than pterostigma; Rs smoothly curved, distal trace fading gradually towards costal margin.

Metasoma: Four exposed terga, internalized sclerites partly exposed, composing a very slender and delicate metasomal tube; sterna convex and undivided medially.

Male: Same as female, except: mandible elongated, flattened, broadly foliaceous, lacking subapical or distal teeth, apparently truncate apically; lower medial margin of clypeus evenly convex; inner margin of antennal sockets touching; scape and pedicel comparatively shorter; flagellum cylindrical, without ventral flattened areas; frons with medial longitudinal groove, extending from median ocellus towards scapal basin; metabasitarsus without two rows of spines; T1–T4 bordered laterally by confluent strong carinae, delimiting the laterotergites; distal elements of genitalia partly exposed, with typical long, slender, and undifferentiated paramere, volsella, and aedeagus.

Etymology

The name is a combination of the words ‘Cenomanian’ (geological age); and - sega, from Amisega Cameron. The name is feminine.

† Cenomanisega krombeini sp. nov.

( Figs 1–3)

ZooBank registration: LSID: urn:lsid:zoobank.org:act:

Description

Holotype female ( DZUP Bur-283). Body length, ~ 4.1 mm.

Head: LID very narrow, ~0.3 × eye width; upper limit of eye surpassing lateral ocellus, inner ocular margins strongly converging toward the upper frons and diverging near posterior margin of head; ocelli deeply inserted, with well-marked small sulci around them, lateral ocelli nearly touching inner ocular margin, closer together than with median ocellus; inner margin of antennal sockets nearly touching; pedicel about three-quarters of F1 length; F1 longer than combined length of F2 + F3; F2 and F3 distinctly shortened, F11 acuminate, slightly longer than F10; subantennal distance ~1 × MOD, clypeus thickened, lower medial margin slightly concave; head finely and sparsely punctate, becoming denser below on face and gena; setation nearly indistinct, short and sparse, becoming slightly longer and denser below on face and occiput; eye glabrous.

Mesosoma: Lower posterior corner of propleuron smoothly round; basisternum not enlarged, disc slightly depressed, posterior process short and broadly round, lateral process long and acute; anterior portion of pronotum short, ~0.3 × total length after transverse groove in mid-line, lateral depression smoothly excavated; profemur thoroughly cylindrical and elongated; metabasitarsus with two rows of short, suberect, thickened spines; dorsum finely and regularly punctate, interspaces slightly rugose, particularly on pronotum and scutum; propodeum mostly smooth, with scarce marginal punctures; metapleuron– propodeum with large medial polished band; dorsum nearly glabrous; some short, suberect, gold setation, becoming denser and longer on legs and mesopleuron; wing membrane entirely setose.

Metasoma: Metasomal terga finely punctate, becoming denser marginally on T3; laterotergites of T1–T2 polished; with short, suberect, gold setation, becoming denser marginally on metasomal sclerites.

Male: See above for sexually dimorphic characters provided in the description of the genus.

Etymology

The species is named after Dr Karl V. Krombein, whose contributions to the biology and systematics of the Amiseginae established the modern foundations for studies involving these fascinating wasps.

Type material

Holotype female in amber piece DZUP Bur-283. The specimen is well preserved and complete. Syninclusion: a nematoceran mosquito.

Paratypes: Two males in the same amber piece, DZUP Bur-505, syninclusion: a wasp ; one female, DZUP Bur-1019 ; one female, DZUP Bur-410, syninclusions: a beetle, five dipterans, and a cricket ; one female, DZUP Bur-577 ; one female, DZUP Bur-771 ; one female, DZUP Bur-074 ; one female, DZUP Bur-1683; and one female, DZUP Bur-1574, syninclusions: four beetles and a wasp .

Additional examined material (non-type)

One male in amber piece DZUP Bur-1772 ; one female, DZUP Bur-1367; and one male, DZUP Bur-1249 .

Remarks

The new genus does not resemble any extant genus of Amiseginae , but it is superficially similar to † Protamisega Evans (known only from the male holotype of Taimyr amber, Russia; Late Santonian, 85.8–84.9 Mya). It can be distinguished from † Protamisega by the combination of the following characters: malar space crossed by longitudinal malar sulcus (lacking in † Protamisega ); pronotum crossed anteriorly and posteriorly by transverse grooves (crossed anteriorly by transverse carina that curves towards midline in † Protamisega ); notaulus deeply impressed, reaching anterior margin of scutum (notaulus evanescent anteriorly in † Protamisega ); parapsidal line well marked (parapsidal line absent or indistinct in † Protamisega ); anterior band of scutellum simple, lacking transverse groove (anterior band of scutellum with transverse groove medially, divided by longitudinal carina in † Protamisega ); dorsum of propodeum moderately long (dorsum of propodeum short, shelf-like in † Protamisega ), lacking longitudinal carinae delimiting a series of foveae (with several longitudinal carinae delimiting a series of foveae in † Protamisega ), posterolateral corner round, lacking posterior propodeal projection (posterolateral corner obtusely angulate in † Protamisega ); posterior declivity of propodeum without carinae radiating outwards (posterior declivity of propodeum with several carinae radiating outwards in † Protamisega ); and dentate tarsal claw (simple in † Protamisega ).