Anoplodactylus gullukensis, Koçak, 2024

Anoplodactylus gullukensis View in CoL n. sp. ( Figures 2–7)

Material examined: (1) One male holotype ( ESFM-PYC/1990-1 ) from Güllük Gulf, 37°15′12″N, 27°37′04″E, collected by Dr. A. Sukatar on 11.09.1990 at depths ranging from 0 to 1 m in brown algae, Cystoseira mediterranea Sauvageau. (2) GoogleMaps One female paratype ( ESFM-PYC/1990-2 ) with the same data as the holotype. (3) GoogleMaps One immature female paratype ( ESFM-PYC/1990-2 ) with the same data as the holotype GoogleMaps .

Diagnosis: An Anoplodactylus species with angular proboscis tip, ventral median suture in ventral view between ventral proboscis antimeres. Trunk without tubercles or setae, no segmentation suture between segments 3 and 4. Immovable finger of cheliphore with 4 spines-like teeth and two spiniform setae on the inner side.

KOÇAK / Turk J Zool

Oviger six-articled. Two cement gland pores without any protruding ducts on the dorsal face of each femur; all leg articles armed with few short setae, propodal sole without cutting lamina, heel bearing two robust spines and one to two slender spines; absence of genital spurs on coxae 2 of leg3 and 4; main claw long-about as long as sole, auxiliary claws minute.

Holotype description: Habitus small size. First trunk segment largest in width and length, without tubercles or setae. Segmentation sutures present between trunk segments 1 and 2, as well as between 2 and 3; no suture between segments 3 and 4 ( Figures 2 and 3A).

Neck short. Ocular tubercule dome shaped, low and wide, not apical cone, close to anterior edge of cephalon, lateral sense organs inconspicuous (this term is sometimes used instead of lateral papilla, e.g., Lehmann et al., 2017, Brenneis, 2022), four eyes well pigmented ( Figure 3A).

Abdomen short, longer than ocular tubercule, almost erect, with three short distal setae, its distal margin is slightly longer than the end of the fourth lateral process ( Figure 3A).

Proboscis cylindrical, slender, somewhat bulging ventrally in lateral view, with two slightly swoollen in about medially, slightly constricted near distal end, about half length of trunk, slightly longer than the length of segments 2 and 3 combined but shorter than segments 1 and 2 combined, in ventral view tip with two protruding angular projections of ventral antimeres (like Anoplodactylus angulatus ), long median suture apparent in ventral view between ventral antimeres ( Figures 3B and 3C).

Oviger of six articles, article 3 longest, article 4 second longest, article 6 triangular with two short setae (one of them well curved), article 5 about half length of article 3, article 1 glabrous, a trace of such a pseudo-articulation in the article 3 (as in A. sandromagnI Krapp, 1996 ), article 2 and 3 with four setae, article 4 with two lateral setae, article 6 about half length of article 5, article 5 with two curved spines and five setae -one of them well curved- ( Figures 3 D-3F).

Cheliphore with a long and slender scape, distally increasing in diameter, overhanging proboscis, slightly curved, sparselysetose.Scapeslightlylongerthanproboscis. Chela shorter than scape. Palm as long as fingers. Fingers subequal, immovable finger almost straight and with 4 spine-like teeth and two spiniform setae on the inner side, movable finger recurved, slightly shorter than immovable finger and with 4 spines-like teeth (the more distal spine reduced to a denticulation) and armed with a spiniform seta on the outer side ( Figures 3G and 3H).

Legs comparatively short, all leg articles armed with few short setae, femur longest, coxa 1 shortest, coxa 2 expanded in diameter distally, mostly so at about half length and slightly longer than coxa 3 ( Figures 4A and 4B). Coxae 2 on legs 3 and 4 without genital spur, only genital pores ventrally on coxae 2 of the legs ( Figure 4A). Tibia 1 slightly longer than tibia 2, and femur slightly longer than tibia 1, the usual subterminal robust setae present near the

KOÇAK / Turk J Zool end of femur and both of tibia ( Figures 4A and 4B). Tarsus short, bearing 6–7 setae ventrally and one seta dorsally. Propodus about as long as tibia 2, curved. Heel distinct, bearing two robust spines and one to two slender spines. Sole almost straight, armed with a row of characteristically curved robust spines with about six to ten, no trace of cutting lamina. Main claw robust, about as long as sole, slightly curved distally. Auxiliary claws minute (less than half length of proximal main claw diameter) ( Figures 4C and 4D).Two cement gland pores without any protruding ducts on the dorsal face of each femur ( Figures 4A and 4B,

Figures 5A and 5B).

Holotype measurements (mm):

Total length from tip of proboscis to tip of the fourth lateral process (ventral): 1.04

Trunk length from frontal margin of cephalic segment to tip of fourth lateral process: 0.85

Trunk width across first lateral process: 0.57

Proboscis length (ventral): 0.42

Abdomen length: 0.12

Cheliphore length: 0.45

Ocular length: 0.08

Oviger lengths: article 1, 0.11; article 2, 0.17; article 3, 0.26; article 4, 0.19; article 5, 0.13; article 6, 0.07

Fourth leg lengths: coxa 1, 0.15; coxa 2, 0.26; coxa 3, 0.20; femur, 0.56; tibia 1, 0.52; tibia 2, 0.49; tarsus, 0.09; propodus, 0.42; main claw, 0.27; auxiliary claw, 0.02

Female: Slightly smaller, but otherwise in close agreement with male ( Figure 6). Femora swollen, with eggs in different size. Genital pores ventrally on coxae 2 of all legs, larger than those in male ( Figure 7A). Main claw curved proximally. Propodal sole bearing more numerous curved robust spines ( Figure 7B). Proboscis as in male, with ventrolateral angular projections at tip corners ( Figure 7C).

Distribution: So far, A. gullukensis n. sp. is known only from the type-locality, Güllük Gulf, Aegean Sea, at a depth of 0 to 1 m.

KOÇAK / Turk J Zool

Etymology: This species is named after the geographical area (Güllük Gulf, Aegean Sea) where the type specimens were collected.