Oedignatha tongbiguan, Zhang & Mu & Zhang, 2025

Oedignatha tongbiguan sp. nov.

Figs 1–4 View FIGURE 1 View FIGURE 2 View FIGURE 3 View FIGURE 4

Type material. Holotype ♂ (MHBUARA-2024-882-17), CHINA: Yunnan Province: Yingjiang County, Tongbiguan Town ( 24.6680°N, 97.5990°E; 913.4 m a.s.l.), 5.X.2024, leg. K. Yu, Y. Ni and Y. Xu. GoogleMaps Paratypes: 3♀ (MHBUARA-2024-882-18–20), same data as holotype GoogleMaps .

Etymology. The specific epithet is derived from the name of the type locality, noun in apposition.

Diagnosis. Males of O. tongbiguan sp. nov. resemble O. dian Lu & Li, 2023 in having a similar shaped embolus and a distally sclerotized conductor, but can be distinguished from the latter by: 1) the retrolateral tibial apophysis bifurcated basally ( Figs 2C View FIGURE 2 , 4C View FIGURE 4 ; vs. apically bifurcated in O. dian , see Lu et al. 2023: figs 5C, 6C); and 2) embolus originating at 8-o’clock position ventrally ( Figs 2B View FIGURE 2 , 4B View FIGURE 4 ; vs. at 9-o’clock position in O. dian , see Lu et al. 2023: figs 5B, 6B). Females are most similar to the females of O. shaanxi Li & Yao, 2023 in possessing a membranous epigynal window and spherical spermathecae, but differs by: 1) the width of the epigynal window about 1.5 times its height ( Figs 3A View FIGURE 3 , 4D View FIGURE 4 ; vs. width about 3 times its height in O. shaanxi , see Chu et al. 2023a: fig. 2A); 2) copulatory ducts narrow ( Figs 3B View FIGURE 3 , 4E View FIGURE 4 ; vs. wide in O. shaanxi , see Chu et al. 2023a: fig. 2B).

Description. Male ( holotype, Figs 1A–B View FIGURE 1 ). Total length 6.55, carapace 3.21 long, 2.09 wide; abdomen 3.34 long, 1.86 wide. Carapace oval, dark brown. Cervical and radial grooves indistinct. Clypeal hump arched ( Fig. 1A View FIGURE 1 ). Labium, sternum and endite reddish brown ( Fig. 1B View FIGURE 1 ). Chelicerae with three pro- and six retromarginal teeth. Eye sizes and interdistances:AME 0.13, ALE 0.14, PME 0.10, PLE 0.11; AME–AME 0.13, AME–ALE 0.17, ALE–ALE 0.70, PME–PME 0.28, PME–PLE 0.29, PLE–PLE 0.97, ALE–PLE 0.11. Legs brown. Leg measurements: Ⅰ 9.41 (2.51, 0.89, 2.35, 2.24, 1.42), II 7.97 (2.25, 0.89, 1.80, 1.89, 1.14), III 7.04 (1.99, 0.81, 1.42, 1.80, 1.02), IV 10.16 (2.69, 0.97, 2.27, 2.81, 1.42); leg formula 4123. Tibial and metatarsal ventral spines depicted in Table 1. Abdomen oval, with complete dorsal scutum ( Fig. 1A View FIGURE 1 ).

Palp ( Figs 2 View FIGURE 2 , 4A–C View FIGURE 4 ). Cymbium with narrow groove basally. Retrolateral tibial apophysis bifurcated at base, prolateral branch S-shaped in ventral view, less sclerotized than retrolateral branch, retrolateral branch triangular in retrolateral view ( Figs 2C View FIGURE 2 , 4C View FIGURE 4 ). Bulb less than half length of cymbium ( Figs 2B View FIGURE 2 , 4B View FIGURE 4 ). Embolus filamentous, originating from 8-o’clock position and extending clockwise to anterior side of conductor ( Figs 2B View FIGURE 2 , 4B View FIGURE 4 ). Conductor widely triangular, with heavily sclerotized edge ( Figs 2 View FIGURE 2 , 4A–C View FIGURE 4 ).

Female ( paratype, Figs 1C–D View FIGURE 1 ) (MHBUARA-2024-882-18). Same as male except for the following: total length 7.19, carapace 2.92 long, 2.25 wide; abdomen 4.27 long, 2.52 wide. Chelicerae with eight retromarginal teeth. Eye sizes and interdistances: AME 0.13, ALE 0.15, PME 0.10, PLE 0.11; AME–AME 0.12, AME–ALE 0.16, ALE– ALE 0.71, PME–PME 0.29, PME–PLE 0.25, PLE–PLE 0.97, ALE–PLE 0.10. Leg measurements: Ⅰ 8.91 (2.41, 0.83, 2.19, 2.14, 1.34), II 7.61 (2.09, 0.82, 1.74, 1.75, 1.21), III 6.57 (1.80, 0.79, 1.29, 1.68, 1.01), IV 9.96 (2.60, 0.85, 2.20, 2.80, 1.51). Tibial and metatarsal ventral spines depicted in Table 1.

Genitalia ( Figs 3 View FIGURE 3 , 4D–E View FIGURE 4 ). Epigynal plate sclerotized, with posteriorly located membranous epigynal window ( Figs 3A View FIGURE 3 , 4D View FIGURE 4 ). Copulatory ducts short, partially visible in dorsal view. Spermathecae spherical. Bursae spherical, clearly smaller than spermathecae. Fertilization ducts pointing anterolaterally ( Figs 3B View FIGURE 3 , 4E View FIGURE 4 ).

Variation. Female (n=3): body length 6.87–7.32.

Distribution. Known only from the type locality.