Haroldiellus woodruffi Skelley and Keller, 2022

Haroldiellus woodruffi Skelley and Keller , new species

Figures 3, 5 View Figures 1–5 , 8–9 View Figures 6–9 , 12–17 View Figures 12–16 View Figure 17

Diagnosis. Length 5.3–5.8 mm, width 2.4–2.7 mm. A member of Haroldiellus distinguished from other species by: Elytral intervals convex, striae deeply impressed; interval color pale with distinct diagonal black marking ( Fig. 3 View Figures 1–5 ). Female lacking median frontal tubercle ( Fig. 8 View Figures 6–9 ); male median frontal tubercle transverse and bifid ( Fig. 9 View Figures 6–9 ). Protibia with dorsal surface lacking punctures ( Fig. 5 View Figures 1–5 ).

Description. Male holotype. Length 5.6 mm, width 2.6 mm. Body robust, not flattened. Body ( Fig. 3 View Figures 1–5 , 12–13 View Figures 12–16 ) color dark reddish brown except head and lateral pronotum pale reddish brown; elytra yellowish brown, striae darkened over disk, intervals with an irregular diagonal dark mark notably contrasting with pale part of intervals; legs paler yellowish brown. Dorsal surface glossy; glabrous; body not fimbriate; ventral surface with moderately long, moderately dense pubescence. Anterior clypeal margin with angles rounded, margin between angles thickened, emarginate. Clypeus and epistome coarsely punctured, flat. Frontal suture trituberculate ( Fig. 9 View Figures 6–9 ), median tubercle broad and bifid. Genae moderately developed, slightly deflexed. Pronotum shiny; with intermixed moderate and fine punctures, all punctures separated by 1–2 times their diameter on disc. Scutellar shield weakly pentagonal, not depressed. Elytra with striae deeply impressed, coarsely punctured; intervals convex, glossy, finely punctured. Ventral surface and metafemur with few coarse punctures. Male metaventrite not deeply excavated, without patch or fringe of dense setae. Protibia lacking dorsal punctures ( Fig. 5 View Figures 1–5 ), unmodified; protibial spur and protarsus unmodified. Inferior mesotibial spur straight at apex, lacking tooth. Metatrochanter with a single seta. Metafemur robust, wider than metacoxa; posterior margin of metafemur smooth, glabrous. Metatibia robust, apically flared, not compressed; apex of metatibia fringed with short, equal spinules. Metatarsus as long as metatibia, tarsomeres slender, cylindrical, with basal tarsomere as long as superior tibial spur, slightly longer than tarsomeres 2 and 3 combined. Epipharynx transverse, side rounded, anterior margin concave towards middle; epitorma conical; corypha slightly protruding, with apical clump of long, slender celtes; pedia pubescent, with several stout, scattered chaetae; chaetopariae moderately dense, moderately elongate ( Fig. 14 View Figures 12–16 ). Male genitalia unmodified; parameres slightly widening at apex, without membranous appendage ( Fig. 15–16 View Figures 12–16 ).

Female allotype. Length 5.3 mm, width 2.4 mm. Body more elongate, less robust than male; median frontal tubercle reduced to simple swelling ( Fig. 8 View Figures 6–9 ).

Variation. Paratypes show some variation in the thickness of the black elytral markings, but they are always well-defined.

Type material. Holotype male ( FSCA) label data: GUATEMALA: El Progresso;/ Sierra de las Minas; nr.Pinalon,/ nr.Finca la Tormenta,/ nr. 15.07222, -89.94891;/ 15-17-V-2010; ~ 2199m;/ P.Skelley, moist oak forest // [red paper] HOLOTYPE ♂ / Haroldiellus / woodruffi P.Skelley / & O.Keller. Allotype female ( FSCA) label data: // GUATEMALA:Guat./ Puerta Parada/ 14°33′24″ -90°27′48″/ 26.III-6.IV.2005 / J.Schuster, 1840m // [blue paper] ALLOTYPE ♀ / Haroldiellus / woodruffi P.Skelley / & O.Keller. Paratype (18 total) label data: COSTA RICA: Prov. of/ San Jose, San Isidro/ de Coronado, Finca, Holanda, 5,500 Ft. // Aphodius / near sallei Har. / # 29 [red ink] Gordon #21 [pencil]/ DET. R.E.Woodruff -72 ( 1m FSCA); GUATEMALA: Guatemala,/ Santa Catarina Pinula ,/ Puerta Parada, Casa-Jack/ Schuster, 1860m,/ N 14.45572 W 90.4653/ 4-11-VII-2009, J.Schuster ( 1m FSCA); GUATEMALA: San Marcos:/ Parque Municipal Refugio/ del Quetzal 1810m 14.93916 / -91.87295 4-11-VI-2015 ZHFalin/ ex. flight intercept trap. cloud/ forest GUAT1F15 099 // Barcode/ SEMC1464610/ KUNHM-ENT (1f SEMC); PANAMA: Chiriqui / Mt. Totumca Lodge/ 19-25-VI-2012 / J.B.Heppner, at light( 1m FSCA); PANAMA: Chiriqui / Prov. 2km W Cerro/ Punta 8°51′N / 82°36′W 3.vi.1977 / H.& A.Howden ( 1m CMN); PANAMA, Chiriqui / Prov. 2-3km E Cerro/ Punta, 2000-2200m, 1./ VI.77 H. & A. Howden // Haroldiellus / n.sp./ det. P. Skelley/ det., 20 2012 ( 1m CMN); Chiquihuite dint./ 15°05′41.0″N / 092°06′04.3″W / 23.VI.2007 - m 2060 // MEXICO / Edo. Chiapas / Leg. Dellacasa M./ & Fresi C. ( 1m, 7f DCGI, 1f FSCA); MEXICO: Chiapas / Mpio: Motozintla/ Boquerón/ 10/V/ 2003 / B. Gómez (1f DCGI); MEXICO: Chiapas / Mpio: Motozintla/ Boquerón/ 29/I/ 2004 / B. Gómez ( 1m DCGI); // MEXICO: Chiapas / Mpio: Motozintla/ Boquerón/ 11/V/ 2003 / B. Gómez ( 1m DCGI);

Distribution. ( Fig. 17 View Figure 17 ). Mexico ( Chiapas), Guatemala ( El Progreso, Guatemala, San Marcos), Costa Rica ( San José), Panama ( Chiriqui).

Etymology. This species is named after Robert Woodruff, friend and mentor, who first studied this species and recognized it as something different, but near H. sallei .

Remarks. Haroldiellus woodruffi , like H. lansbergei , appears to be a species occupying higher elevations.