Diamesa hamaticornis Kieffer, 1924

Diamesa hamaticornis Kieffer View in CoL

( Figs 1 View FIGURES 1–7 , 10 View FIGURES 8–14 , 23–24 View FIGURES 23–26 )

Diamesa hamaticornis Kieffer, 1924: 56 View in CoL ; Pagast 1947: 485; Wülker 1959: 350; Serra-Tosio 1971: 236; Schmid 1993: 37; Langton & Visser 2003: 23; Ashe & O’Connor 2009: 276; Makarchenko et al. 2022a: 78.

Material examined. 3 adult males, RUSSIA: Republic of North Ossetia-Alania, Irafsky District, Caucasus Mountains , Terek River basin, Karaugomidon River , alt. 1476 m a. s. l., 04.I.2018, 42.8942 N 43.6973, leg. D. Palatov ; 2 adult males, the same data, except Alagirsky District, Caucasus Mountains, Terek River basin, Skazdon River , alt. 1943 m a. s. l., 27.VII.2019, 42.7842 N 43.9037 E, leg. D. Palatov GoogleMaps ; 2 adult males, Magadan Region, Olsky District, mouth of Ola River , 03. V.2019, 59.580514 N, 151.272686 E, leg. E. Khamenkova GoogleMaps ; 1 adult males, Republic of Khakassia, Abakan City, valley of the Abakan River , poplar forest, alt. 245 m a. s. l., 06.IV.2020, 53.713750 N, 91.522111 E, leg. S. Dragan GoogleMaps ; 2 adult males, the same data, except Republic of Khakassia, Askizsky District, Askiz River , from surface of snow, 05.XI.2020, 53.339417 N, 89.888500 E, leg. S. Dragan GoogleMaps ; 1 adult male, KYRGYZSTAN: Talas Region, Chychkan Кiver Valley, Chychkan River, approximately 10–15 km below the pass from the Suusamyr Мalley, alt. 2454 m a. s. l., 6.II.2016, 42.19895 N, 72.989667 E, leg. D. Palatov GoogleMaps ; 2 adult males, MONGOLIA: Govi- Altay Aymag, Uliastein Gol River at the confluence of Baga Gol River , alt. 1989 m a. s. l., 22.IX.2019, 45.484600 N 94.277067 E, leg. D. Palatov. GoogleMaps

Description

Adult male (n = 3, except when otherwise stated). Total length 3.7–3.9 mm. Total length/wing length 1.02–1.22. Coloration. Dark brown to brown. Legs brown. Wings grayish to gray, venation brownish.

Head. Eyes hairy, reniform. Temporal setae including 4 preoculars, 11 verticals. Clypeus with 8 setae. Antenna with 13 flagellomeres ( Fig. 1 View FIGURES 1–7 ) and little reduced plume of setae, 131–560 µm long; terminal flagellomere with 1–2 subapical setae, 36–52 μm long; AR 0.50–0.67. Palpomere length (in μm): 48–52, 88–96, 120–140, 124–140, 180–216. Palpomere 3 in distal part with sensilla capitata with diameter 12–16 μm. Head width/palpal length 0.99– 1.08.

Thorax. Antepronotum with 10–11 ventrolateral setae. Dorsocentrals 9–15, prealars 6–10. Scutellum with 15– 38 setae.

Wing. Length 3.2–3.64 mm, width 1.0– 1.12 mm. Anal lobe rounded to rounded-angular. Squama with 22–30 setae, 72–88 μm long. R + R 1 with 22–27 setae, R 4+5 with 10–11. RM/MCu 2.7. Costa extension 98 μm long.

Legs. Spur of front tibia 54 µm long. Spurs of mid tibia 48 µm long. Spurs of hind tibia 84 and 56 µm long. Hind tibial comb with 18 setae. Length (μm) and proportions of leg segments are as in Table 4.

Hypopygium ( Figs 23–24 View FIGURES 23–26 ). Tergite IX with 11–15 setae from one side and anal point, 176–180 µm long. Laterosternite IX with 8–13 setae. Transverse sternapodeme 180–240 μm long and 52–60 μm wide. Gonocoxite 360 µm long; inferior volsella 128–140 µm long, wide in basal 3/4, narrow and finger-shaped in short distal quarter, with long setae along margin ( Fig. 23 View FIGURES 23–26 ). Basimedial setal cluster with 12–17 setae, 112–196 µm long ( Fig. 24 View FIGURES 23–26 ). Gonostylus 320 µm long, widened in basal half and narrowed in the distal, a long inward margin with pale short setae, apically with megaseta, 8 µm long. HR= 1.13.

Pupa. Described by Serra-Tosio (1971) and Langton & Visser (2003).

Larva. Described by Schmid (1993).

Distribution. Palaearctic species. Widely distributed in the Western Palaearctic ( Ashe & O’Connor, 2009) and known also in East Siberia ( Makarchenko et al. 2022a) and Magadan Region of the Russian Far East.