Thalictrum przewalskii Maximowicz (1877: 305)

Thalictrum przewalskii Maximowicz (1877: 305) View in CoL . Figs. 1‒4 View FIGURE 1 View FIGURE 2 View FIGURE 3 View FIGURE 4 .

Type:— CHINA. Gansu: Regio silvestris declivit., borealis jugi a fl. Tetung meridium versus, ad silvarum margines, 27 July/8 August

1872, N. M. Przewalski 348 [holotype LE 01014048!; isotype LE 01014049!; see Grubov (2000) and Zeng et al. (2021a)]. = T. sexnervisepalum Wang (2017a: 679) , syn. nov. Type:— CHINA. Sichuan: Chongzhou city, Anzihe Nature Reserve, Jiguanshan, in grass, alt. 3100 m, 27 July 2007, Z. B. Feng, D. H. Zhu

& X. J. Li 4214 [holotype PE01962228 !; isotypes WCSBG (2 sheets, barcodes unavailable)!]. Fig. 1 View FIGURE 1 . = T. lasiogynum Wang (2017b: 414) . Type:— CHINA. Sichuan: Pingwu county, Wanglang Nature Reserve , Baishagou , on roadside, 3110 m, 19 July 2008, L. M. Lu 2008337

(holotype PE 02108941!). = T. latistylum Wang (2017a: 676) . Type:— CHINA. Gansu: Wenxian county, Baishuijiang Nature Reserve, Qiujiaba, on grassy slope, 2900 m, 27 June 2006, Baishuijiang

Exped. 454 (holotype PE 02108940!). = T. rockii Boivin (1945: 115) .

Type:— CHINA. Gansu: Upper Tebbu country (= Tewo county), below Shimen, Tsaluku valley, in forests of spruces, willows, etc., 11000 ft., July‒August 1925, J. F. Rock 13054 (holotype GH 00038614!).

Description:—Perennial herbs.Roots fibrous.Stem 50–120cm tall, more or less puberulous in young plants, particularly so on the upper part, later usually glabrescent but sometimes still puberulous on nodes. Basal leaves withered at anthesis. Cauline leaves 3–4-ternate; leaf blade triangular, 7–25 cm long and broad; leaflets obovate, rhombic-elliptic, ovate or suborbicular, 0.5–3 cm long, 0.5–2.5 cm broad, herbaceous, adaxially sparsely puberulous, later often glabrescent, abaxially often puberulous, base broadly cuneate, rounded or subcordate, apex obtuse or acute, 3-lobed; lobes obtusely 2‒3-dentate, apex obtuse; veins submerged adaxially, prominently raised abaxially; petioles to 6 cm long, with stipules at base, sparsely puberulous, later glabrescent; stipules oblong, sheath-like, membranous, margin entire. Inflorescence a terminal thyrse; rachis sparsely or densely puberulous, later glabrescent. Pedicels 3‒10 mm long, sparsely or densely puberulous, later glabrescent. Flowers bisexual, erect. Sepals 4‒5, caducous, ellliptic, obovate or nearly oblong, 2.5–5 mm long, 1.5‒2.2 mm broad, adaxially glabrous, abaxially totally or distally puberulous or ciliate only on the upper margin, white or tinged with yellowish-green. Stamens 30‒50, 5‒8 mm long; filaments 4‒7 mm long, proximally filiform, distally thicker, white; anthers ca. 1 mm long, oblong, apex obtuse or acute, pale yellow. Carpels 4–9, more or less puberulous, stipitate; stipe 0.3‒1 mm long; ovary obliquely obovate, compressed; style 1.5‒1.8 mm long, straight; stigma oblong, ca. 0.5 mm long. Achenes 8‒15 mm long, more or less puberulous when young, glabrescent or sparsely puberulous when mature, stipitate, drooping; stipe 1.5‒7 mm long, filiform; body obliquely broadly obovate, ca. 6 mm long, ca. 5 mm broad, compressed, shallowly 2‒3-ribbed; style persistent, straight.

Phenology:—Flowering from June to August; fruiting from July to September.

Distribution and habitat:— Thalictrum przewalskii is widely distributed in Beijing, Chongqing, Gansu, Hebei, Henan, Hubei, Hunan, Nei Mongol, Ningxia, Qinghai, Shaanxi, Shanxi, Sichuan, and Xizang, China ( Fig. 5 View FIGURE 5 ). It grows in bushes, forests or on grassy slopes at elevations of 750‒3500 m above sea level.

Additional specimens examined:— CHINA. Beijing: Fengtai, C.G. Yang 982 (IBK, KUN); Mentougou, W.Y. Hsia 2074 (NAS), E. Licent 3120 (BM, K, P), Beijing Chin. Trad. Med. Exped. 417 (PE), Q.P. Xiang & S. Qu GPS243- 16 (PE), Y. Yabe s.n. (NAS), C.G. Yang 76 (PE), C.G. Yang 303 (PE), C.G. Yang 1385 (PE), C.G. Yang 1687 (NAS, PE); Miyun, J.W. Wang s.n. (PEY); Pinggu, Pinggu Exped. 572 (PE); Yanqing, W. Tong 2006071018 (PEM), J.W. Wang s.n. (PEY); Precise locality unknown, G. Hemeling 186 (E). Chongqing: Wuxi, K.H. Yang 58742 (IBSC, PE, SZ), K.H. Yang 58947 (CDBI, PE); Precise locality unknown, P.G. Farges s.n. (K, P), P. Pelliot & L. Vaillant 865 (P).

Gansu: Hezuo, S.Q. Guo & P. Du 169 (HNWP), S.Q. Guo & P. Du 174 (HNWP); Huixian, C.Y. Chang 833 (IBSC, WUK), P.C. Kuo 4920 (WUK); Jingchuan, C.P. Wang 17073 (PE); Jingtai, Y.P. Hsu & C.Y. Yu 3171 (WUK); Jone, Anonymous 8 (PE), Anonymous D0035 (HNWP), R.C. Ching 956 (E), Sino-Brit. Qinghai Alp. Gard. Soc. Exped. 122 (E), T.J. Tong & M. Tang 91 (IBSC), C.P. Wang 5340 (PE); Kangle, G.Y. Yang 92-002 (KUN); Lanzhou, Anonymous s.n. (PE), E. Licent 4442 (P); Linxia, X.G. Sun et al. 2096 (PE), X.G. Sun et al. 2098 (PE), X.G. Sun et al. 2117 (PE), X.G. Sun et al. 2474 (PE), X.G. Sun et al. 20010519 (PE); Luqu, Taohe Exped. 216 (KUN); Maqu, G.L. Zhu 1785 (PE); Minxian, K.S. Hao 651 (PE), W. Purdom s.n. (K); Sunan, Y.C. Ho 3550 (WUK); Tewo, J.F. Rock 12635 (NAS); Tianshui, K.T. Fu 14161 (WUK), K.T. Fu 16114 (KUN, SZ, WUK), W.Y. Hsia 5700 (PE), W.Y. Hsia 5710 (WUK), W.Y. Hsia 5715 (PE); Tianzhu, Anonymous 10 (WUK), T.N. Ho 2270 (HNWP), Y.C. Ho 4512 (PE, WUK), Y.C. Ho 4821 (PE, WUK), R.F. Huang 2607 (HNWP), R.F. Huang 2675 (HNWP), Loess-Plateau Exped. 5076 (WUK); Weiyuan, Yellow River Exped. 1607 (NAS, PE, WUK); Wenxian, C.Y. Chang 6897 (HNWP, WUK), Y.P. Zeng 289 (IBSC), Y.P. Zeng et al. 115 (IBSC); Xiahe, K.T. Fu 1228 (IBSC, PE, WUK), W. Qi 374 (HNWP), C.P. Wang 6892 (PE, WUK), C.P. Wang 6897 (PE, WUK), C.P. Wang 7328 (IBSC, PE, WUK), J.S. Yang 91-1043 (KUN, PE), L.H. Zhou & H.Z. Zhang 1437 (HNWP); Yongdeng, T.N. Ho 2041 (HNWP), R.F. Huang 2372 (HNWP), Loess-Plateau Exped. 4738 (WUK), J.P. Luo & L. Wang 245 (IBSC), X.G. Sun et al. 6012 (PE), X.G. Sun et al. 6245 (PE); Yuzhong, Y.C. Ho 1803 (WUK), Y.C. Ho 5950 (PE, WUK), G. Ho & C.P. Wang 126 (WUK), B. Lei 76 (PE), Yellow River Exped. 3353 (PE); Zhugqu, P.C. Kuo 5126 (WUK), P.C. Kuo 5387 (WUK), C.P. Wang 14414 (PE, WUK), Q.E. Yang & H.Z. Kong 98-36 (PE), Q.E. Yang & H.Z. Kong 98-38 (PE); Precise locality unknown, Anonymous 68 (PE), G.N. Potanin s.n. (K), N.M. Przewalski s.n. (E, LE, PE), J.F. Rock 12835 (E). Hebei: Chengde, Chengde Exped. 1547 (PE), Herbarium Team 2142 (WUK), J.W. Wang & S.P. Kong 57 (PEY), Yu s.n. (PEY), W.X. Zhang et al. 56 (PE); Chicheng, Anonymous 7294 (HNWP, PE); Fuping, Fuping Exped. 98 (PE); Huailai, Anonymous 1749 (PE), Anonymous 2024 (HNWP, PE), X.L. Huang et al. 1652 (PE); Laishui, K.M. Liou 2282 (LBG, NAS, PE), K.M. Liou 2533 (PE); Laiyuan, Anonymous 4-2306 (HNWP, PE); Pingquan, M.D. Chen & T.Y. Sun 26 (PE); Pingshan, J.W. Wang & S.P. Kong 738237 (PEY); Qinglong, Z.T. Wang et al. 140258 (PE); Xinglong, Herbarium Team 2142 (PE), T.N. Liou & P.Y. Fu 4769 (NAS, PE), Miyun Exped. 502 (PE), T. Tang 2064 (PE, WUK), W.T. Wang 2694 (PE), Yu 170 (PE, PEY), Yu 198 (PE); Yuxian, Anonymous

576 (BJM), Anonymous 1992 (PE), Anonymous 5030 (HNWP, PE), J.X. Duan 421 (PE), Hebei Exped. 498 (PE), Hebei Exped. 1749 (PE), Herbarium Team 2381 (PE, WUK), X.L. Huang et al. 1387 (HNWP, PE), X.L. Huang et al. 1463 (HNWP, PE), H.W. Kung 1227 (E, PE), C.F. Li 11927 (PE), C.F. Li 61527 (PE), W. Limpricht 2575 (K), Y. Liou 11316 (PE, PEY), Y. Liou 11535 (IBSC, PE), C.Y. Wu & C.Y. Yang 2686 (PEY), C.Y. Wu & C.Y. Yang 2915 (PE, PEY), C.Y. Wu & C.Y. Yang 36921 (PE), C.Y. Wu & C.Y. Yang 36922a (PEY), C.Y. Wu & C.Y. Yang 37134 (PE), C.Y. Wu & C.Y. Yang 37279 (PEY), Xiaowutaishan Exped. 74223 (PE), C.W. Wang 61512 (PE), C.W. Wang 61527 (PE), C.W. Wang 61692 (IBSC, PE, PEY), C.Y. Yang 36948 (PEY); Zhuolu, J.X. Duan 206 (PE), W.Y. Hsia 2400 (E, PE, SZ), C.W. Wang 60643 (PE), C.W. Wang 61005 (PE), Yang 7014 (PE, PEY), C.G. Yang 76 (PE), C.G. Yang 303 (PE), C.G. Yang 471 (KUN, PE, WUK), C.G. Yang 982 (PE), C.G. Yang 1172 (PE), C.G. Yang 1573 (NAS, PE); Zunhua, H.F. Chow 40848 (PE), K.M. Liou 412 (PE), H.T. Tsai 50031 (IBSC), H.T. Tsai 50315 (IBK, IBSC), Tungling Exped. 136 (KUN, PE), Tungling Exped. 151 (PE); Precise locality unknown, H.F. Chow 65 (IBK), F. Hugh s.n. (BM), T.T. Li 228 (PE), Y. Liou 11871 (IBSC). Henan: Lingbao, J.M. Li et al. 140805102 (HEAC), J.M. Li et al. 14080562 (HEAC); Luanchuan, M.H. Jin 24 (HENU), Loess-Plateau Exped. 1784 (WUK), Loess-Plateau Exped. 1953 (WUK); Lushi, K.M. Liou 5002 (PE), K.M. Liou 5053 (PE), J.P. Luo & Y. Zou 35 (IBSC), J.P. Luo & Y. Zou 36 (IBSC), Q.E. Yang et al. 4184 (IBSC); Songxian, Anonymous 22248 (NAS), K.C. Kuan & T.L. Tai 2132 (NAS, PE), Y.P. Zeng & Q.L. Huang 437 (IBSC); Yiyang, Anonymous 569 (NAS). Hubei: Shennongjia, 1980 Sino-Amer. Exped. 159 (E, KUN, NAS), Hubei Plant Exped. 25128 (PE), B.B. Liu 2280 (PE), C.L. Zhang 874 (IBSC); Xingshan, Z.D. Chen & X.T. Ma 961273 (PE); Precise locality unknown, E.H. Wilson 2334 (K). Hunan: Shimen, L.H. Liou 17810 (NAS). Nei Mongol: Harqin, Y. Sun 1064 (HIMC); Liangcheng, Y.Q. Ma 53 (HIMC); Ningcheng, S.X. Li 4540 (IFP), J.W. Wang s.n. (HIMC). Ningxia: Guyuan, C.P. Wang 13677 (PE, WUK); Jingyuan, C.P. Wang 17073 (WUK); Longde, C.P. Wang 13145 (IBSC, KUN, WUK), C.P. Wang 13167 (WUK). Qinghai: Baima, S.L. Chen et al. 341 (HNWP), Sino-Brit. Qinghai Alp. Gard. Soc. Exped. 822 (E), Z.D. Wei 180 (WUK), L. Wang, X.Q. Guo & Y.P. Zeng 3578 (IBSC), Y.H. Wu 26220 (HNWP), Y.H. Wu 26241 (HNWP); Datong, C.W. Chang 2498 (HNWP, WUK), H.B. Chen 87055 (PEM), H.B. Chen 87063 (PEM), P.C. Kuo & W.Y. Wang 12278 (HNWP), B.W. Li 72-126A (HNWP), Y.L. Li 659 (HNWP), K.M. Liou 5994 (IBSC, PE, WUK), Loess-Plateau Exped. 5277 (WUK), L.J. Mei 193 (HNWP), Y.H. Wu 16074 (HNWP), Y.H. Wu 16118 (HNWP), Y.H. Wu 16136 (HNWP), Y.H. Wu 16175 (HNWP), Y.H. Wu 16178 (HNWP), Y.H. Wu 19766 (HNWP), Y.H. Wu 19790 (HNWP), Y.H. Wu 19850 (HNWP), Y.H. Wu 20180 (HNWP), Y.H. Wu 23433 (HNWP), Y.H. Wu et al. 17322 (HNWP), Y.H. Wu et al. 17322 (HNWP), Y.H. Wu et al. 17381 (HNWP), Y.H. Wu et al. 17387 (HNWP), Y.H. Wu et al. 17432B (HNWP), Y.H. Wu et al. 17441 (HNWP), Y.H. Wu et al. 17450 (HNWP), Y.H. Wu et al. 17512 (HNWP), Y.H. Wu et al. 17527 (HNWP), S.Z. Yan 701 (WUK), Z.H. Zhang 4354 (PE), Z.H. Zhang et al. 4364 (HNWP), Z.H. Zhang et al. 4365 (HNWP); Golog, W.Y. Wang et al. 26857 (HNWP), W.Y. Wang et al. 27311 (HNWP), W.Y. Wang et al. 27477 (HNWP), W.Y. Wang et al. 27613 (HNWP), W.Y. Wang et al. 27626 (HNWP); Gonghe, Y.H. Wu 9712 (HNWP), Y.H. Wu 9715 (HNWP); Guide, T.N. Ho 606 (PE), T.N. Ho 1171 (HNWP, PE), S.W. Liu 3278 (HNWP, PE), Y.H. Wu 187763 (HNWP); Haiyan, Anonymous 1713 (HNWP), P.C. Kuo & W.Y. Wang 12278 (HNWP, PE), S.W. Liu 1898 (HNWP), Y.H. Wu et al. 7497 (HNWP); Henan, Sino-Brit. Qinghai Exped. 1997 750 (E); Huangyuan, Anonymous 53 (WUK), G.X. Liu & Y.Y. Tian s.n. (PE), Y.H. Wu 9155 (HNWP), Y.H. Wu 9169 (HNWP); Huangzhong, S.W. Liu 2053 (HNWP), S.W. Liu 2119 (HNWP), Sino-Brit. Qinghai Exped. 1997 1189 (E), S.X. Wang 1116 (HNWP), Y.H. Wu et al. 17592 (HNWP), Y.H. Wu et al. 17610 (HNWP), Y.H. Wu et al. 17662 (HNWP), Y.H. Wu et al. 17666 (HNWP), Y.H. Wu et al. 17820 (HNWP), Y.H. Wu et al. 17827 (HNWP), Y.H. Wu et al. 17835 (HNWP), Y.H. Wu et al. 18996 (HNWP); Huzhu, T.N. Ho 1575 (HNWP), R.F. Huang 1913 (HNWP), P.C. Kuo 25573 (HNWP), Y.H. Wu 2678 (HNWP), Y.H. Wu 19282 (HNWP), Y.H. Wu 19301 (HNWP), Y.H. Wu 19323 (HNWP), Y.H. Wu 19344 (HNWP), Y.H. Wu 38615 (HNWP), Y.H. Wu 38626 (HNWP), Y.H. Wu 38644 (HNWP), Y.H. Wu 38652 (HNWP), Y.H. Wu 38673 (HNWP); Jainca, Y.H. Wu & L.J. Mei 16605 (HNWP), Y.H. Wu & L.J. Mei 16650 (HNWP), Y.H. Wu & L.J. Mei 16748 (HNWP); Jigzhi, Golog Exped. 595 (HNWP), Xizang Trad. Med. Exped. 880 (HNWP); Ledu, X.F. Lu et al. 361 (HNWP), X.F. Lu et al. 433 (HNWP), Z.H. Wu & Yan . S. Chen 26 (WUK), Y.H. Wu 15569 (HNWP), Y.H. Wu et al. 3297 (HNWP), Y.H. Wu et al. 3298 (HNWP), Y.H. Wu et al. 3470 (HNWP, PE), Y.H. Wu et al. 3662 (HNWP, PE), Y.H. Wu et al. 3668B (PE), Y.H. Wu et al. 8272 (HNWP), Y.H. Wu et al. 8353 (HNWP), Y.H. Wu et al. 8397 (HNWP), Y.H. Wu et al. 8457 (HNWP); Madoi, Y.H. Wu et al. 5187 (HNWP); Maqen , Anonymous 127 (HNWP), B.Q. Ma 2127 (HNWP), Maqen Exped. 429 (HNWP), Sino-Brit. Qinghai Exped. 1997 460 (E), Sino-Brit. Qinghai Exped. 1997 509 (E), Y.H. Wu 18757 (HNWP), Y.H. Wu 18761 (HNWP), Y.H. Wu 18780 (HNWP), Y.H. Wu 18789 (HNWP), Y.H. Wu 18793 (HNWP), Y.H. Wu 20670 (HNWP), Y.H. Wu 20713 (HNWP), Y.H. Wu et al. 4613 (HNWP), Y.H. Wu et al. 5591 (HNWP), X.Y. Wu & X.L. Zhang QH2014012 (BNU); Menyuan, P.C. Kuo 7316 (HNWP, KUN), P.C. Kuo & W.Y. Wang 12123 (HNWP, PE), Loess-Plateau Exped. 5005 (WUK), Qinghai-Gansu Exped. 2551 (PE, WUK), Qinghai-Gansu Exped. 2682 (PE, WUK), Y.H. Wu 23123 (HNWP), Y.H. Wu 23129 (HNWP), Y.H. Wu 23133 (HNWP); Minhe, T.N. Ho 606 (HNWP), S.W. Liu 2731 (HNWP, PE), Y.H. Wu 24422 (HNWP), Y.H. Wu 24548 (HNWP), Y.H. Wu 24818 (HNWP), Y.H. Wu 24891 (HNWP), Y.H. Wu 25050 (HNWP), Y.H. Wu 25161 (HNWP), L.H. Zhou 2388 (HNWP), L.H. Zhou 2562 (HNWP); Ping'an, X.F. Lu et al. 99 (HNWP); Qilian, K. Rong 2078 (WUK); Tongde, Anonymous 33 (HNWP), T.N. Ho et al. 137 (HNWP), J.F. Rock 14271 (E, K, NAS), B. Tian & M.F. Ji 2010-QH-031 (HNWP), Y.H. Wu et al. 4696 (HNWP), Y.H. Wu et al. 4809 (HNWP), Y.H. Wu et al. 5229 (HNWP), Y.H. Wu et al. 5953 (HNWP), Y.H. Wu et al. 6212 (HNWP), Y.H. Wu et al. 6840 (PE), Y.H. Wu et al. 7317 (HNWP); Tongren, S.W. Liu & D.S. Luo 2-1378 (HNWP), Loess-Plateau Exped. 5718 (WUK); Ulan, Anonymous s.n. (PE); Xining, H.F. Bidley s.n. (BM), H.F. Bidley 5 (K), K.S. Hao 790 (PE, WUK), Y.H. Wu et al. 2206 (HNWP), Z.H. Zhang et al. 487 (HNWP), Z.H. Zhang et al. 2011 (HNWP), Z.H. Zhang et al. 4282 (PE); Xunhua, C.Y. Chang 19134 (WUK), P.C. Kuo & W.Y. Wang 25208 (HNWP), Loess-Plateau Exped. 5772 (WUK), G.B. Shi 3 (IBSC); Yushu, T.N. Ho et al. 2013 (BM, E, HNWP, PE), Z.D. Wei 22438 (HNWP), B.Q. Xu et al. Xianh0332 (IBSC); Zekog, T.N. Ho 526 (HNWP), P.C. Kuo 25874 (HNWP), P.C. Kuo 25931 (HNWP), P.C. Kuo 25937 (HNWP), P.C. Kuo 26084 (HNWP), L.H. Zhou & L.N. Sun 1655 (HNWP); Precise locality unknown, J.F. Rock 14125 (E). Shaanxi: Baoji, K.M. Liou 10922 (WUK); Chang'an, S.F. Li et al. 11009 (XBGH); Fengxian, K.M. Liou 10703 (PE, WUK); Foping, P.C. Kuo 1727 (IBSC, PE, WUK); Huyi, K.T. Fu 14320 (WUK), K.T. Fu 14377 (HNWP, WUK), WUK Exped. 2022 (WUK); Langao, Yan S. Chen et al. 4494 (WUK); Luonan, C.P. Wang 15829 (KUN, PE, WUK); Meixian, BNU Qinling Exped. TB401 (BNU), K.T. Fu 8349 (KUN, WUK), G. Giraldi s.n. (K, P), Y.H. Guo & Z.X. Hu 460 (IBSC, WUK), F. Hugh s.n. (BM), S.F. Li et al. 12119 (XBGH), Y.M. Liang 220 (WUK), W. Purdom s.n. (K), W. Purdom 419 (K), Qinling Exped. 349 (WUK), Qinling Exped. 433 (WUK), Qinling Exped. 495 (WUK), P.C. Tsoong 2279 (WUK), Z.P. Wei 1193 (WUK), Z.P. Wei 1345 (WUK), J.Y. Wu et al. 20 (PE), J.Y. Wu et al. 45 (PE), J.Y. Wu et al. 64 (PE), Zhu et al. 1352 (KUN, PE); Mianxian, Anonymous 25 (HNWP), Anonymous 2-021 (HNWP); Ningshan, K.T. Fu 17926 (IBK, WUK), H.W. Kung 3012 (PE), X.Y. Liu 15117 (WUK); Pingli, Z.H. Wu 1295 (WUK), G.Y. Xu 4446 (WUK), Y.P. Zeng & Q.L. Huang 420 (IBSC); Taibai, K.T. Fu 9221 (WUK), K.T. Fu 9361 (WUK), K.T. Fu 10359 (WUK), T.N. Liou & P.C. Tsoong 938 (PE), T.N. Liou & P.C. Tsoong 2204 (WUK), Zhu et al. 2547 (KUN, PE); Zhenba, K.T. Fu 11576 (WUK); Zhenping, Anonymous 297 (WUK), T.S. Ying et al. 626 (WUK); Zhouzhi, Qinling Plant Res. Exped. 399 (NAS). Shanxi: Fanzhi, C.H. Duan 178 (PE), Shanxi Exped. 603 (PE); Huozhou, Yellow River Exped. 722 (KUN, NAS); Jiaocheng, Anonymous 7-52 (SXU), Anonymous 3-051 (SXU), Anonymous 1508021 (SXTCM), Anonymous 1515025 (SXTCM), Anonymous 1518042 (SXTCM), Anonymous 1521005 (SXTCM), Anonymous 151061013 (SXTCM), Q.Y. Deng 201607017 (SXTCM), S. Gao 201609034 (SXTCM), W.Y. Hsia 1126 (PE), W.Y. Hsia 1225 (PE), X. Li 201608047 (SXTCM), S.Z. Liu 1622123 (SXTCM), X.Y. Liu 21284 (HNWP, PE), X.Y. Liu 21555 (HNWP), Loess-Plateau Exped. 2951 (WUK), X.L. Ma 1613090 (SXTCM), S.N. Wang 1615079 (SXTCM), K. Wu 1624117 (SXTCM), C.X. Xu 1605037 (SXTCM), X.Y. Yu 94739 (SXPHC), J.Y. Zhang 1606049 (SXTCM), Z. Zhang 1623109 (SXTCM), Z. Zhang 1623146 (SXTCM); Jiexiu, K.M. Liou 1242 (PE); Lingshi, Anonymous 851077 (SXPHC); Lishan, Yellow River Exped. 2417 (PE, WUK), Yellow River Exped. 2518 (PE, WUK); Ningwu, K.M. Liou 1609 (PE), K.M. Liou 1737 (PE), X.Y. Liu 15117 (HNWP, WUK), X.Y. Liu 15948 (HNWP, WUK), X.T. Zhang 4025 (PE); Qinxian, K.C. Kuan & Y.L. Chen 1057 (HNWP), K.C. Kuan & Y.L. Chen 1059 (PE); Qinyuan, Shanxi Exped. 2642 (WUK); Shanyin, Y.W. Tsui 2334 (HIMC); Wutai, K.C. Kuan & Y.L. Chen 1549 (HNWP, PE); Wuzhai, Yellow River Exped. 2911 (NAS, PE); Xingxian, Yellow River Exped. 2559 (PE, WUK); Yuanqu, H. Smith 6648 (BM, PE), Loess-Plant. Exped. 2265 (WUK); Precise locality unknown, J.F. Rock 14125 (E). Sichuan: Aba, Golog Exped. 771 (HNWP), S. Jiang et al. 1198 (PE), K.Y. Lang, L.Q. Li & Y. Fei 2046 (KUN, PE); Baiyu, D.E. Boufford et al. 37147 (KUN, PE), D.E. Boufford et al. 37157 (PE); Barkam, C.Y. Chang & H.F. Chow 23716 (PE, WUK), W.P. Fang et al. 22506 (PE), K.Y. Lang et al. 2187 (KUN, PE), H. Li 70614 (IBSC), H. Li 71545 (IBSC, PE, SZ), H. Li 74480 (IBSC, KUN, NAS, PE, SZ, WUK), C.L. Wu 32049 (PE), C.L. Wu 32507 (NAS, PE), L. Xie 200109015 (PE); Chongzhou, Y.P. Zeng & X.X. Zhou 1297 (IBSC); Dawu, Anonymous 95 (SZ), Anonymous 4242 (CDBI), S. Jiang et al. 9666 (PE), Q.E. Yang, L. Wang & X.Q. Guo 4014 (IBSC), Q.E. Yang, L. Wang & X.Q. Guo 4018 (IBSC); Dege, Anonymous 4227 (CDBI), Anonymous 7487 (CDBI, PE), D.E. Boufford et al. 36521 (KUN, PE), D.E. Boufford et al. 36540 (KUN, PE), D.E. Boufford et al. 36801 (P, PE), S.X. Jia 2343 (PE), Y.W. Tsui 4406 (PE), Y.W. Tsui 4469a (PE), Y.H. Wu 30148 (HNWP), Q.E. Yang & Q. Yuan 456 (IBSC), Xizang Suppl. Exped. 6105 (KUN); Garze, D.E. Boufford et al. 33995 (KUN, PE), K.N. Wang 1718 (PE); Heishui, G.N. Potanin s.n. (PE), H. Li 73229 (NAS, PE); Hongyuan, C.Y. Chang & H.F. Chow 22365 (KUN, NAS, PE, SZ), C.Y. Chang & H.F. Chow 22506 (NAS, SZ), C.Y. Chang & H.F. Chow 23716 (KUN, NAS, SZ, WUK), S.Y. Chen 5715 (NAS, SZ), X.J. He et al. 133504 (SZ), X.J. He et al. 133505 (SZ), X.J. He et al. 133569 (SZ), X.J. He et al. 143760 (SZ), X.J. He et al. 146403 (SZ), X.J. He et al. 146705 (SZ), X.J. He et al. 148067 (SZ), X.J. He et al. 148089 (SZ), X.J. He et al. 148190 (SZ), X.J. He et al. 148967 (SZ), H. Li 74565 (E, IBSC, NAS, PE, SZ), H. Li 74637 (IBSC, KUN, NAS, PE, SZ, WUK), H. Li 74649 (IBSC, NAS, PE, SZ), H. Li 74698 (IBSC, KUN, NAS, PE, SZ, WUK), B.C. Ni et al. 376 (CDBI), Sichuan Veg. Exped. 9192 (CDBI, IBSC, PE), Q.S. Zhao 93 (BM, E, PE, SZ), Q.S. Zhao 254 (BM, E, K, P, PE); Jinchuan, P.X. li 10424 (PE), P.X. li 10541 (PE); Jiuzhaigou, Anonymous 604 (SM), Y. Shi et al. 159 (HNWP), Q.X. Li & X.C. Zhao 3113 (PE), M. Tamura et al. 805 (CDBI), L. Wang et al. 3622 (IBSC); Kangding, K.C. Kuan & W.T. Wang 839 (K, PE); Lixian, S.Y. Chen et al. 5460 (SZ), S.Y. Chen et al. 5464 (NAS, SM), C. Ho & T.L. Chow 13643 (IBSC, LBG, NAS, PE), W.K. Hu et al. 46186 (IBSC), H. Li 74160 (PE), H. Li 74162 (IBSC, NAS, SZ), T.P. Soong 46186 (SZ), H.Q. Yang 72 (SZ); Luhuo, Y.T. Chang & K.Y. Lang 103 (PE); Maoxian, C. Ho & T.L. Chow 13643 (SZ); Pingwu, Z.Y. Chen s.n. (SZ), X.J. He et al. 129299 (SZ), X.J. He et al. 129342 (SZ), L. Wang 1413 (IBSC), L. Wang 1442 (IBSC), Y.P. Zeng & Q.L. Huang 338 (IBSC), Y.P. Zeng & Q.L. Huang 446 (IBSC); Sertar, D.E. Boufford et al. 34637 (KUN, PE), Z.S. Qin 6531 (CDBI, PE); Serxu, Anonymous 209 (SM), Q.E. Yang et al. 4059 (IBSC); Songpan, D.E. Boufford et al. 40435 (PE), G. Heerebout s.n. (L), H. Smith 2526 (BM), H. Smith 2692 (PE), M. Tamura et al. 717 (CDBI), C.P. Wang 7795 (IBSC, PE, WUK), H.C. Young 72 (SZ), P. Zhuang et al. 2004138 (WSCBG); Wenchuan, X.J. He et al. 128443 (SZ), X.J. He et al. 150288 (SZ); Xiaojin, S.S. Chang & Y.X. Ren 6139 (CDBI, PE, SZ); Xinlong, Y.P. Zeng & Q.L. Huang 401 (IBSC); Zamtang, D.E. Boufford et al. 39068 (PE), Y.H. Wu 30901 (HNWP), Y.H. Wu 30910B (HNWP), Y.H. Wu 30933 (HNWP), Y.H. Wu 30948 (HNWP); Zoige, Anonymous 122 (SM), Anonymous 170 (SM), Anonymous 424 (SM), C. Ho 20564 (SM), S. Jiang et al. 1418 (PE, SZ), K.Y. Lang et al. 1977 (KUN, PE), Sichuan Trad. Med. Plant Exped. 20564 (NAS), Q.E. Yang et al. 4110 (IBSC), X. Yin et al. 214 (HNWP), D.H. Zhu et al. 4577 (PE, WSCBG); Precise locality unknown, K.L. Chu 7309 (IBSC, NAS), G.N. Potanin s.n. (P). Xizang: Chagyab, X.X. Huang & Z.B. Zhong 16291 (IBSC); Jomda, D.E. Boufford et al. 41974 (KUN, PE), Y.T. Chang & K.Y. Lang 163 (PE), Inst. Biol. Xizang Exped. 2399 (HNWP), Qinghai-Xizang Exped. 12457 (HNWP, KUN, PE), L. Wang et al. 3429 (IBSC).