Macroscytus reflexus SIGNORET, 1883

Macroscytus reflexus SIGNORET

(Figs 279­282)

Macroscytus reflexus SIGNORET, 1883a: 466 ; LETHIERRY & SEVERIN 1893: 71.

Macroscytus acutus SIGNORET, 1883a: 469 ; LETHIERRY & SEVERIN 1893: 70; JEANNEL 1913: 15; LINNAVUORI 1993: 25, syn. n.

Macroscytus excavatus SIGNORET, 1883a: 471 ; LETHIERRY & SEVERIN 1893: 70; MANCINI 1953: 203, 1954: 157, 1956: 95; WAGNER 1963: 91; MEDLER 1980: 122; LINNAVUORI 1977: 40, 1980: 35, 1993: 22; LIS 1991d: 176, syn. n.

Macroscytus inermipes HORVÁTH, 1904: 253 ; BERGROTH 1908: 149; HORVÁTH 1919: 237; LINNAVUORI 1977: 40 (syn. with M. excavatus by LINNAVUORI 1993: 22), syn. n.

DESCRIPTION

Body 9.25­11.90 mm in length, 5.04­6.59 mm in width, from yellowish brown to almost black, sometimes corium slightly paler than pronotum and scutellum.

Head dorsally at first sight looking impunctate, but bearing hardly visible tiny punctures, lateral parts of paraclypei wrinkled; clypeus subapically without setigerous punctures, each paraclypeus submarginally with 2­4 setigerous puncture bearing long hair­like setae (Fig. 279); ocular index 2.10­2.64; interocellar index 4.0­8.5; length of antennal segments (in mm): 0.34­0.63: 0.59­0.76: 0.73­ 0.85: 0.90­1.13: 0.94­1.15.

Puncturation of pronotal disc more or less dense, punctures small, clearly visible; calli, umbones and posterior fourth of pronotal disc impunctate; discal transverse impression behind calli shallow, developed only laterally, usually acompanied by a row of coarse punctures; lateral margins of pronotum with 6­8 submarginal setigerous punctures bearing long hair­like setae (Fig. 279). Anterior convexity of propleuron apically punctate with small punctures, posterior convexity distinctly anteriorly punctate with large punctures, propleural depression with a band of coarse punctures and wrinkles.

Scutellar disc punctate with almost evenly spaced punctures.

Corium densely punctate; clavus with one complete and two partial rows of punctures; mesocorial disc densely and almost evenly punctate; exocorium densely punctate; costa flattened, basally expanded, separated from exocorium only in its basal half, costal margin with 2 setigerous punctures bearing long hair­like setae (Fig. 279).

Male mid femur with a row of small teeth; male hind femur with a row of well visible teeth and denticles on margins, subbasal and subapical teeth large (variability of edentation was presented by LINNAVUORI, 1993), male hind tibia with a row of numerous denticles; female hind femur with small (sometimes indistinct) subapical tooth, female tibia simple.

Abdominal sterna more or less polished, punctate only around spiracles and trichobothria.

Male genitalia as in Figs 280­282.

TYPE DATA

Syntype (s) male (s) of Macroscytus reflexus SIGNORET : South Africa (lost or destroyed) . Neotype male (present designation) of Macroscytus reflexus SIGNORET : South Africa, Natal, Empangeni area, 28°45̕S 31°55”E, 20­II­ 86, 110m, P. Reavell, light ( NMP) .

Lectotype female (designated by LINNAVUORI 1993: 25) of Macroscytus acutus SIGNORET : Kenya, Mombasa ( BMNH) . Lectotype male (designated by LINNAVUORI 1993: 22, as Holotype) of Macroscytus excavatus SIGNORET : East Africa, Lac Hyarsa [= Nyassa Lake ] ( BMNH) . Lectotype male (designated by LINNAVUORI 1977: 41) of Macroscytus inermipes HORVÁTH : Sao Thomé ( HNHM) .

MATERIAL EXAMINED

Neotype of Macroscytus reflexus , Lectotype of Macroscytus acutus , Lectotype of Macroscytus excavatus , Lectotype of Macroscytus inermipes , and 229 other specimens: Angola: 1 ex. ( MNHN) . Burkina Faso: Ouagadougou , 1 ex. ( ZIP) . Cameroon: Nkolbisson , 3 exx. ( MNHN, UO) ; Yaoundé , 1 ex. ( MNHN) . Central African Republic: La Maboke , 16 exx. ( MNHN, UO) ; Boukoko , 3 exx. ( MNHN) . Congo: Bouenza , 1 ex. ( MNHN) ; Brazzaville , 85 exx. ( HNHM, MNHN, UO) ; Sibiti , 1 ex. ( MNHN) ; Odzala , 1 ex. ( MNHN) ; Dimonika, Mayumbe , 7 exx. ( MNHN) . Eritrea: Ghinda , 2 exx. ( NNML, ZIP) . Ethiopia: Bahor Dar , 10 exx. ( SMNS, UO) ; Gidole, Prov. Gamu Gofa , 5 exx. ( SMNS, UO) ; Wollo­Ataye , 1 ex. ( MNHN) ; Kébré­Mengist , 1 ex. ( MNHN) ; Lake Langano, Shoa , 3 exx. ( ZSMC) . Gabon: Ovenda near Libreville , 1 ex. ( ZIZM) . Guinea: Mt. Nimba, Plateau de Zouguepo , 1 ex. ( MNHN) . Ivory Coast: Lamto , 1 ex. ( MNHN) . Kenya: Malindi , 3 exx. ( MNHU) ; Mombasa , 2 exx. ( MNHU, SMNS) ; Mt. Elgon National Park, near Chapnyalil Cave , 1 ex. ( HNHM) ; Ukunda, Diani Beach , 1 ex. ( HNHM) ; Tsavo West National Park , 1 ex. ( HNHM) . Liberia: 1 ex. ( ZIZM) . Mozambique: 1 ex. ( ZSMC) ; Beira , 1 ex. ( MNHN) ; Lourenco Marques, Matola , Vila Salazar, 1 ex. ( JEC) . Namibia: Kavango, Popa­Falls , 6 exx. ( MNHU, MNHU, UO) ; East Caprivi, Katima Mulilo , 3 exx. ( MNHU, UO) . Ruanda: Kissenyi , 1 ex. ( MNHN) . Senegal: Kedougou, 2 exx. ( MNHN) . Sierra Leone: Makeni , 12 exx. ( MZLU, UO) . South Africa: Natal, Empangeni area, 14 exx. ( NMP, UO) ; Natal, Kube Yini Game Reserve , 3 exx. ( NMP, UO) ; Natal, Mkuzi Game Reserve , 7 exx. ( MNHU, NMP, UO) . St. Thomé Island: Bombaim , 1 ex. ( USMB) . Tanzania: Arusha distr., Naberera env., 3 exx. ( USMB, UO) ; Morogoro , 4 exx. ( HNHM, UO) . Togo: Lama Kara , 1 ex. ( ZIZM) . Uganda: Nyabyeya , 3 exx. ( MNHU, UO) . Zambia: Chisamba nr Lusaka, 1 ex. ( NMP) ; Kafue National Park, Chunga Camp , 1 ex. ( MNHU) ; 5 km S Choma , 1 ex. ( MNHU) . Zimbabwe: Matabeleland, Matopos National Park , 3 exx. ( MNHU, UO) ; Zimbabwe Ruins , 1 ex. ( NMP) ; Kariba, 5 exx. ( JEC, UO) .

DISTRIBUTION

Angola, Burkina Faso (first record), Cameroon, Central African Republic,?Chad, Congo, Eritrea, Ethiopia, Gabon (first record), Gambia, Ghana, Guinea (first record), Ivory Coast, Kenya, Liberia (first record), Mozambique (first record), Namibia (first record), Nigeria, Ruanda (first record), Senegal (first record), Sierra Leone, South Africa, St. Thomé Island, Sudan, Tanzania, Togo (first record), Uganda, Zaire, Zambia, Zimbabwe.

REMARK

The type material of Macroscytus reflexus from South Africa was not found in the collections containing SIGNORET ̕s specimens (it was apparently lost or destroyed). Since the other three species ( Macroscytus acutus , M. excavatus , M. inermipes ) are found to be conspecific with the former, it was necessary to designate a neotype for M. reflexus .