Anemone hupehensis (Lemonine) Lemonine

Anemone hupehensis (Lemonine) Lemonine View in CoL in Lemonine & Fils (1910: 40). A. japonica var. hupehensis Lemonine in

Lemonine & Fils (1908: 42). Eriocapitella hupehensis (Lemonine) Christenhusz & Byng in Christenhusz et al. (2018:

75), syn. nov. Figs. 1–4 View FIGURE 1 View FIGURE 2 View FIGURE 3 View FIGURE 4 , 7–9 View FIGURE 7 View FIGURE 8 View FIGURE 9 , 13D View FIGURE 13 .

Type:— CHINA. Hupeh (= Hubei): no type designated ( Wang, 2021; see below) .

= Anemone robustostylosa Miau (1993: 57) View in CoL , syn. nov.

Type:— CHINA. Guangxi: Leye, Yachang forest farm, alt. 950, roadside, 30 August 1989, Y. C. Zhong 890811 (holotype SYS 00053030!). Fig. 1 View FIGURE 1 .

= Anemone discoideogyna Wang (2021: 103) View in CoL , syn. nov.

Type:— CHINA. Sichuan: Shimian, Jinwo xiang, alt. 2179 m, in scrub, 18 July 2018, X. Y. Zhu et al. G 10222 (holotype PE 02347292!, isotypes PE 02347293!, PE 02347294!). Fig. 2 View FIGURE 2 .

Description:—Perennial herbs, (20–) 30–120 cm tall. Rhizome branched, oblique or erect, 8–12 cm long, 0.5–0.7 cm in diameter, woody. Basal leaves 3–5, usually ternate, sometimes 1 or 2 or even all simple, long petiolate; petiole 5–35 cm long, sparsely or densely pubescent, base short-sheathing; central leaflet long petiolulate (petiolule 1–6.5 cm long), blade ovate or broadly ovate, 4–11 cm long, 3–10 cm broad, apex acute or acuminate, base rounded or cordate, undivided or 3–5-lobate, margin serrate, sparsely strigose on both surfaces; lateral leaflets similar to central one but smaller. Scape erect, sparsely pubescent; cyme 2- or 3-branched, many-flowered, occasionally unbranched, only 3- flowered. Involucral bracts 3, slightly unequally sized, petiolate; petiole 0.5–6 cm long, base sheathing; bract blade similar to that of leaves, ternate, 3–7 cm long. Pedicel 3–10 cm long, sparsely or densely pubescent, lateral ones with small, paired bracteoles. Sepals 5 (flower single) or ca. 20 (flower double, in cultivated plants), purple, purple-red, pink or white, obovate, 20–30 mm long, 13–20 mm broad, abaxially velutinous, basal veins 5–9, vein anastomoses more than 10. Stamens 4–6 mm long; filament filiform; anther yellow, ellipsoid. Pistils more than 180, stipitate, ca. 1.5 mm long; ovary velutinous; style glabrous; stigma rectangular. Achene body ovoid, ca. 2 mm long, ca. 1 mm broad, lanate, hairs 3–4 mm long; style straight, short.

Phenology:—Flowering from July to October; fruiting from August to November.

Distribution:— Anemone hupehensis is native in Chongqing, Gansu, Guangxi, Guizhou, Henan, Hubei, Hunan, Jiangxi, Shaanxi, Sichuan, Taiwan, Yunnan, and Zhejiang, China ( Fig. 12 View FIGURE 12 ). It grows in scrub, on grassy slopes or along streamside in hilly regions at altitudes of 400–2600 m. It has been widely cultivated or naturalized in southern China.

Additional specimens examined:— CHINA. Chongqing: Banan, Anonymous 9892 (SM), B.H. Li 1555 (SM); Beibei, Jinyun Exped. 0171 (CQNM); Bishan, Bishan Exped. 427 (SM); Changshou, Changshou Exped. 500115131029- 929LY (IMC); Chengkou, Bashan Exped. 0589 (PE), 0630 (PE), 1494 (PE), 2050 (PE), E. Bodinier 1881 (P), D.P. Ho 106872 (NAS, PE), P.Y. Li 3002 (WUK), 3739 (WUK), 3968 (WUK), 5252 (PE, WUK), 6583 (PE, WUK), 8005 (WUK), T.L. Tai 102050 (PE), 102840 (NAS, PE), 103244 (NAS, PE), 103500 (PE), 103501 (IBSC, NAS, PE), 103838 (IBSC, NAS, PE), 104323 (IBSC, NAS, PE, WUK), 104393 (IBSC, NAS, PE, WUK), 104635 (IBSC, NAS, PE, WUK), 104819 (CDBI, IBK, NAS, PE), 104879 (PE), 105854 (IBK, NAS, PE), 105896 (NAS, PE), 106872 (NAS, PE), 107025 (IBK, NAS, PE), 107076 (IBK, NAS, PE), 107467 (PE); Dianjiang, Dianjiang Exped. 0575 (SM), 500231-131012-715 (IMC); Fengdu, Anonymous 467 (SM), Fengdu Exped. 500230-001-0728 (IMC); Fengjie, Anonymous 0411 (SM), C.Y. Chang 5852 (SM), 25858 (IBSC, KUN, NAS, PE), H.F. Chow 10419 (IBSC, NAS, PE), H.F. Chow & H.Y. Su s.n. (PE), 110419 (NAS, PE, WUK), 110638 (IBSC, NAS, PE, WUK), 110866 (IBSC, NAS, PE, WUK), 111217 (IBSC, NAS, WUK), 111324 (IBSC), M.Y. Fang 23974 (IBSC, PE); Fuling, Anonymous 0499 (SM), Fuling Exped. 500102-001-0766 (IMC); Jiangjin, Jiangjin Exped. 500116-422 (IMC), Jiangjin Group 0267 (SM), Z.Y. Liu et al. S-3038 (IMC), S-3402 (IMC), S-3834 (IMC); Kaizhou, Bashan Exped. 2551 (PE); Nanchuan, Anonymous 214 (WUK), 0928 (SM), 1029 (SM), 1054 (SM), 95215 (NAS), 96891 (IMC, SM), C.H. Hsiung & T.L. Chow 92554 (IBSC), 92624 (IBSC, PE), 93705 (IBSC, SM, WUK), 93741 (IBSC), 95315 (SM), C.Y. Huang 201 (NAS), K.C. Kuan et al. 1055 (PE), 2001 (PE), 2153 (PE), 2322 (PE), G.F. Li 63122 (IBSC, NAS), 63245 (IBSC, NAS), 63949 (IBSC, NAS), 64399 (IBSC, SZ), S.Q. Liu 96598 (SM, IMC), Z.Y. Liu 14153 (IMC, PE), 14510 (PE), Nanchuan Med. Plantation 276 (SM), 1988 (PEM), 2878 (SM), Nanchuan Exped. 5001190455 (IMC), Plant Geog. Exped. 674 (CDBI, PE), M.L. Sheh et al. 6437 (NAS), Q. Yuan LCS11475 (IBSC), X.B. Zhang 2 (NAS); Pengshui, Anonymous 1174 (SM), Z.Y. Liu 1092 (SM); Qianjiang, Anonymous 920 (SM), Z.Y. Liu 1122 (IMC, SM), Qianjiang Exped. 50114-292 (IMC), Z.C. Zhao 88-1401 (PE); Qijiang, Qijiang Exped. 13-500222-LY-269 (IMC); Rongchang, Rongchang Exped. 549 (SM); Shizhu, Y. Chen 3062 (CDBI); Tongliang, Tongliang Exped. 0480 (SM); Wulong, Anonymous 1076 (SM), 1220 (SM), 1251 (SM), X.H. Jiao 0245 (SM), Z.Y. Liu 180038 (IMC, PE), 181309 (PE), W.H. Wang 3482 (CDBI); Wushan, L.Q. Chen & J.Q. Xiong chuanhua-531 (CCAU), H.F. Chow & H.Y. Su 10985 (PE), 109591 (IBSC, WUK), 109857 (NAS, WUK), 109924 (IBSC, PE, WUK), 110137 (IBSC, NAS, WUK), Sanxia Exped. 3351 (PE), K.H. Yang 65498 (CDBI, NAS, WUK); Wuxi, Y.D. Chen et al. 2000 (PE), 2484 (PE), K.L. Chu 1378 (NAS), IBCAS Sanxia Exped. 0120 (PE), 0236 (PE), Long et al. 730-282 (SM), P.L. Peng 0903 (SM), K.H. Yang 5358 (NAS), 59144 (CDBI, IBSC, NAS, PE, WUK), 59265 (CDBI, IBSC, NAS), 65085 (CDBI, IBSC, NAS, PE, WUK), 65358 (CDBI, IBSC, WUK), 65498 (IBSC); Xiushan, Anonymous 1115 (SM), Z.Y. Liu L1197 (IMC, SM), Z.C. Zhao 1851 (PE); Yongchuan, T.H. Tu 120 (PE); Youyang, Anonymous 296 (SM), 1530 (SM), J.A. Wang 287 (CDBI), Y. Wu et al. YY20181005_0010 (JIU), Youyang Exped. 500242-413 (IMC), Z.C. Zhao & J.S. Ma 2853 (CDBI, PE); Yunyang, Yunyang Exped. 500235- 131010-923 (IMC); Zhongxian, Y.T. Hou & C.Y. Guo 20150621105 (QFNU), D.S. Luo 1482 (SM), Sanxia Exped. 2522 (PE), Zhongxian Exped. 500233-131024-926 (IMC). Gansu: Kangxian, Z.Y. Zhang 16743 (PE, WUK), 16949 (PE, WUK); Wenxian, Baishuijiang Exped. 2427 (PE), 2600 (PE), 5611 (PE), Y.C. Ho 911 (PE, WUK), Z.P. Wei 2878 (HHBG, WUK), J.C. Xing 21847 (WUK), J.X. Yang & Z.X. Hu 3507 (IBSC, PE, WUK), 3714 (IBSC, PE, WUK), Z.Y. Zhang 8059 (WUK), 8289 (NHWP, WUK), 8325 (WUK), 8355 (WUK), 9138 (WUK), 9361 (WUK), 10316 (WUK), 10437 (WUK), 10632 (WUK), 10728 (WUK), 11990 (WUK), 12092 (WUK), 12719 (WUK), 12781 (WUK), 12817 (WUK), 12895 (WUK), 13095 (WUK), 13357 (WUK), 13427 (WUK), 13568 (WUK), 13728 (WUK), 13798 (WUK), 13982 (PE, WUK), 14311 (PE, WUK), 14944 (PE, WUK), 14457 (WUK), 14716 (WUK), 14944 (PE, WUK), 15221 (PE, WUK); Zhugqu, Z.P. Wei 2464 (HHBG, WUK). Guangxi: Debao, Debao Exped. 451024151011034LY (IBK), J. Huang & Y.F. Huang 21926 (GXMG), C. Ren & Y.F. Huang 19499 (GXMG); Donglan, F. He et al. 451224181128001LY (GXMG), C. Ren & Y.F. Huang 19461 (GXMG); Du’an, T.N. Huang & Y.K. Lee 62474 (IBK); Gongcheng, J.X. Han 29 (PE), B.N. Zhang 050 (IBK); Huanjiang, Huanjiang Exped. 451226130825010LY (GXMG, IBK), H.Z. Lü & Q.S. Dong 15666 (GXMG), R.C. Peng & J.C. Yang ML0840 (IBK), C. Ren & Y.F. Huang 19446 (GXMG), F.N. Wei 2040 (IBK), F.N. Wei & Y. Liu M0002 (IBK), Yunnan-Guizhou-Guangxi Exped. 70036 (IBK); Jingxi, Y.S. Huang et al. LYJX0360 (GXMG), S.P. Ko 55804 (IBK), H.Z. Lü & B.Y. Huang 16978 (GXMG); Leye, Anonymous 603488 (IBK), J.X. Huang & S.M. Xiong HJX10046 (IBSC), HJX10056 (IBSC), HJX10060 (IBSC), HJX10061 (IBSC), Z.T. Li 603287 (IBK), Y.D. Peng 15995 (GXMG); Lingchuan, Lingchuan Exped. 450323130927057LY (GXMG, IBK); Lingui, Lingui Exped. 450322150818018LY (GXMG), Sino-German Exped. 380 (IBSC); Lingyun, Anonymous 92518 (IBK), Z.T. Li 603204 (IBK), Lingyun Exped. 451027121017011 (GXMI), X.Q. Liu 28490 (IBK); Longsheng, Kwangfu Exped. 00931 (IBK, IBSC), 01070 (IBK, PE, SZ), 1197 (IBK), G.Z. Li 62788 (IBK), B. Liu 491 (PE), Longsheng Exped. 50182 (IBK, NAS), 450328130903032LY (IBK, GXMG), 450328130917034LY (IBK, GXMG), C.H. Lü & Y.Z. Wei 20029 (IBK), H.F. Qin 700818 (IBK, IBSC), R.R. Yang 84092 (PE); Longlin, J.Y. Liang K1473 (IBK); Luocheng, Luocheng Exped. 451225130424033LY (IBK); Napo, C.C. Chang 12331 (IBK, IBSC), 14104 (IBK, IBSC), Y.F. Chen & S.X. Yu 08080 (PE), Z.Y. Li et al. 1818 (PE), H.N. Qin et al. 2603 (PE), H. Wang 6588 (PE), X.S. Yu et al. 278 (GXMG); Nandan, C. Huang 40965 (IBK), Z.T. Li 601557 (IBK), Nandan Exped. 451221171029025LY (GXMG); Quanzhou, Y.C. Chen 301 (GXMI, IBK); Rong’an, Rong’an Exped. 450224190904020LY (IBK); Rongshui, S.H. Chun 15911 (IBK, IBSC), 16058 (IBK, IBSC, LBG, NAS, WUK); Tian’e, Beijing Exped. 893119 (PE), 893196 (PE), 896978 (PE), Tian’e Exped. 451222140816019LY (GXMG); Tianlin, Anonymous 5077 (IBK), Y.K. Lee P00716 (IBK), N.F. Li et al. 17235 (GXMG), N.F. Li & C. Ren 18704 (GXMG), Z.T. Li 600692 (IBK), Tianlin Exped. 451029121020033 (GXMG), 451029130723006 (GXMG); Xing’an, Z.Z. Chen 51298 (IBK, IBSC), C.S. Chung 83446 (IBK), G.Z. Li 11090 (IBK), 11991 (IBK), 12826 (IBK), 14133 (IBK, PE), 15156 (PE), S.L. Yu 900267 (IBK, IBSC); Ziyuan, Z.Z. Chen 52048 (IBK), T.N. Huang 61015 (IBK), Ziyuan Exped. 450329160811017LY (IBK), 450329160817006LY (IBK). Guizhou: Anlong, Guizhou Exped. 3622 (IBSC, PE, WUK), C.F. Lan 150 (GZTM); Anshun, Anshun Exped. 1965 (PE); Bijie, P.H. Yu 190 (GZTM), 444 (IBSC, PE, WUK), 790 (PE, WUK); Ceheng, Z.Y. Cao 532 (HGAS, PE), F.C. Wang 539 (PE); Changshun, M.T. An CS-077 (GZAC); Chishui, Bijie Exped. 1352 (HGAS, PE), Z.Y. Cao & Z.T. Wang 20 (PE), 441 (PE), H.T. Zhao 520381160503016LY (GZTM); Dafang, Bijie Exped. 921 (HGAS, PE), C.L. Liu 182 (GZTM); Dejiang, M.T. An 3653 (PE), 3791 (PE), 3929 (PE), H. Li 522227160607003LY (GZTM), N Guizhou Exped. 1755 (IBK, NAS, PE), 2689 (PE), Z.C. Zhao 88-2589 (PE); Dushan, S Guizhou Exped. 74-990 (HGAS, PE); Duyun, Y.Y. Cao 1-A-3-23 (GNUG), Y.Y. Liu 86-1019 (GNUG), Y.S. Xiao 008 (GZTM), Z. Xu 019 (GZTM); Guanling, Guanling Exped. 530 (GZTM); Guiding, Y. Tsiang 5391 (NAS), R. You & D.X. Ji 31-2 (GZTM); Guiyang, M.T. An 5288 (PE), M.C. Wang 520112131025018LY (GYBG), Wudang Exped. 520112131110271LY (GYBG); Hezhang, Y. He 522428151110142LY (GZTM), P.H. Yu 1368 (LBG, PE, WUK); Huaxi, Z.Y. Cao 346 (PE); Huangping, S.H. He 329 (GZTM), S Guizhou Exped. 2805 (HGAS, NAS, PE); Jiangkou, Z.P. Jian et al. 32144 (HGAS, PE), Y.K. Li 8248 (HGAS, IBSC), M.L. Yang 024 (GNUG), W.X. Zhang et al. 144 (PE); Jinping, Z.C. Long 522628140822179LY (GZTM), Z.R. Wang 522628141118369LY (GZTM); Jinsha, C.H. Huang 5 (PE), 58 (PE); Kaili, Y.T. Chang & T.S. Chang 1159 (HGAS, PE), S Guizhou Exped. 3450 (NAS, PE); Leishan, Q.H. Chen 3318 (HGAS), H. Yang 020241 (GZTM), H.J. Zhan 522634150913012LY (GZTM); Libo, Q.H. Chen 2254 (HGAS), Y.K. Li 10128 (HGAS), Z.Z. Peng 15965 (GXMG), Y.M. Wu 201811077 (QNUN); Liupanshui, D.C. Wang 123 (GZTM); Liuzhi, P. Su 62 (GZTM), 143 (GZTM); Longli, B. Tang 522730150518020LY (GZTM); Meitan, Y.T. Chong 19 (GZTM), Guizhou Med. Exped. 225 (GZTM), N Guizhou Exped. 2511 (HGAS, NAS, PE); Nayong, Bijie Exped. 304 (HGAS, PE); Panxian, Anshun Exped. 1222 (HGAS, PE), C.Z. Dang 804 (HGAS), S.P. Gu 2-232-4 (GZTM), Z.F. Jiang 520222140717001LY (GZTM), Z.X. Pan 104 (GZTM), 2-232-2 (GZTM), X.Y. Tang 658 (GZTM), S.C. Zhang 2-232-4 (GZTM); Pingba, Anshun Exped. 1451 (HGAS, KUN, PE); Pu’an, Anshun Exped. 1323 (HGAS, PE), Y.J. Gao & K.H. Jiang 64 (GZTM), G.Y. Luo 64 (GZTM); Qianxi, Z.X. Pan 270 (GZTM); Qinlong, C.J. Yang 66 (GZTM); Qingzhen, S.Z. He 20422 (GZTM), Sichuan- Guizhou Exped. 1694 (PE); Renhuai, X.L. Wang 2225 (PE); Rongjiang, Z.P. Jian et al. 51478 (PE), S Guizhou Exped. 3040 (HGAS, NAS, PE), S.G. Yang et al. 522632190423728LY (GZTM); Sandu, Anonymous 60 (GZTM); Shibing, Wulingshan Exped. 3610 (PE); Shiqian, Q.H. Chen 94193 (HGAS), Wulingshan Exped. 2994 (PE); Sinan, H.T. Gong 023 (GZTM), S.T. Li 167 (GZTM); Suiyang, Suiyang Exped. 520323140909073LY (GZTM), 520323151224045LY (GZTM); Tongren, Horticulture Group 071 (GNUG), M.L. Yang 024 (GNUG), S.Q. Yang 6 (GZTM); Tongzi, R.B. Zhang ZRB236 (ZY); Wangmo, G.F. Wang 1-0637 (PE), 1-0908 (PE), 1-0909 (PE); Xifeng, M.Z. Yang 810445 (HGAS); Xingren, T.S. Chang & Y.T. Chang 7554 (IBSC, NAS, PE), C.Z. Dang 1416 (HGAS), C.Z. Dang & P. Dang 541 (PE), C.L.Hu 311 (GZTM); Xingyi, Anshun Exped. 650 (PE), C.Y. Deng522301140630359LY (GZTM), 522301140902490LY (GZTM); Xishui, Bijie Exped. 1550 (PE), 1620 (PE), C.W. Wu 1250 (PE); Xiuwen, M.T. An 11 (PE); Yanhe, Y.L. Cao & F.T. Pu 2866 (PE), N Guizhou Exped. 2276 (HGAS, NAS, PE), J.W. Xiao 5885 (PE); Yinjiang, Z.P. Jian et al. 30355 (HGAS, PE), 31254 (HGAS, PE), J.J. Xu 129 (GZTM); Yuping, B. Feng 522223140323016LY (GZTM), L. Wang 522223140325081LY (GZTM), Y.S. Zhang 522223150807042LY (GZTM); Yuqing, A.C. Shi & A.G. Liu 123 (GZTM); Weining, Z.X. Wang & C.J. Li 194 (GZTM), Weining Exped. 522427140610130LY (GZTM); Zhenfeng, S.X. Tan 4 (PE), Q.B. Wu 522325180919339LY (GZTM), G.X. You 93 (GZTM); Zhenning, Zhenning Exped. 80 (GZTM); Zhenyuan, Tozii s.n. (NAS); Zheng’an, Q.H.Chen5301 (HGAS), 5313 (HGAS); Zhijin, A.H.Wang522425150402018LY (GZTM); Ziyun, S.Z. Xu et al. 0307073 (GZTM), Ziyun Exped. 488 (GZTM); Zunyi, S. Li XQS089 (ZY), Sichuan- Guizhou Exped. 957 (PE), 1175 (PE), A.N. Steward et al. 81 (PE), Y.L. Wu 4054-1 (GZTM). Henan: Xichuan, Henan Depart. Forest. 1460 (PE). Hubei: Badong, Q.L. Chen et al. 1878 (NAS, PE), Hubei Shennongjia Plant Exped. 24684 (HIB), R.H. Huang 329 (WUK), M.Z. Qian 1568 (PE, WUK), Sino-Amer. Bot. Exped. 53 (NAS, KUN, PE), D.R. Wan s.n. (PE), Y.M. Wang 178 (IBK, WUK), C.M. Zhao et al. EX2745 (PE), EX3100 (PE), EX3137 (PE); Changyang, C.P. Wang 11428 (PE, WUK); Dangyang, L.Y. Tai & C. H. Chien e1313 (PE); Enshi, F.H. Chen 5005 (HIB), K.H. Fu & T.S. Chang 1456 (NAS, PE), M.X. Nie 1456 (IBSC), F.L. Teng 16110737 (CSFI); Fangxian, Anonymous 0374 (LBG), K.M. Liou 8951 (NAS, PE, WUK), 9084 (PE, WUK), 9304 (NAS, PE, WUK), J.C. Xing 16356 (HIB, WUK), 16976 (HIB, WUK), 17119 (HIB, WUK), 17977 (WUK); Hefeng, H.C. Li 217 (WUK), 6109 (HIB, IBSC, PE, WUK), 6511 (HIB, IBK, IBSC, PE, WUK), 6536 (IBK, IBSC, PE, WUK), Y.M. Wang 6047 (HIB, PE); Jianshi, H.C. Chow 1356 (NAS, PE), L.Y. Tai & C.H. Chien 1313 (NAS), Q. Yuan 2014-1 (IBSC); Jingmen, J.Q. Wu & X.Y. Ma 8147 (HIB); Junxian, Anonymous 0081 (LBG), K.R. Liu 81 (HIB, PE), J.C. Xing 10486 (WUK), 10592 (WUK), 10616 (WUK), 10640 (WUK), 10756 (WUK), 10808 (WUK), 11097 (WUK), 11943 (WUK), 12417 (SZ), 12634 (WUK), 12768 (WUK), 13063 (WUK), 13091 (WUK), 14296 (WUK), 15571 (WUK), 16084 (HIB, WUK), G.F. Zhu 1800 (IBSC); Laifeng, H.C. Li 5369 (IBK, IBSC, PE, WUK), 7197 (HIB, PE, WUK); Lichuan, Anonymous 197 (NAS), K.H. Fu & T.S. Chang 1572 (NAS, PE, WUK), R.H. Huang 3721 (HIB), M.X. Nie 1572 (IBSC), L.Y. Tai & C. H. Chien 1059 (NAS, PE), J.L. Yan 392 (WUK), Z.C. Ye 306 (HIB); Shennongjia, Y.T. Hou et al. 16001 (QFNU), C.M. Hu 684 (HIB, LBG), R.H. Huang 710 (HIB, WUK), Hubei Plant Exped. 11405 (HIB, PE), Hubei Shennongjia Plant Exped. 11386 (HIB, PE), 11953 (HIB, PE), 31480 (HIB, PE), 31942 (HIB, PE), M.Z. Qian 1568 (HIB, WUK), Z.J. Ren 2618 (PE), Shennongjia Exped. 21847 (HIB, PE), Sino-Amer. Bot. Exped. 754 (HIB, NAS), C.Q. Wang IV3030287 (CCAU), S.X. Yang 0391 (IBSC), Zhou & Dong 76026 (PE); Shiyan, C.S. Wang 254 (PE); Wufeng, G.X. Chen & D.G. Zhang SCSB-HC-2008283 (KUN); Wuhan, H.C. Li 1456 (PE); Xianfeng, H.C. Li 7535 (IBSC, PE, WUK), 9043 (IBSC, PE, WUK), 9143 (NAS), 9500 (HIB, IBSC, PE), Y.M. Wang 6471 (PE); Xingshan, Q.H. Liu 310 (HIB, IBK), T.P. Wang 11967 (PE, WUK), C.M. Zhao EX2092 (PE), C.M. Zhao et al. EX1417 (PE); Xuan’en, H.C. Li 5050 (PE), 5087 (HIB), Y.M. Wang 5140 (HIB, PE); Yichang, C.M. Hu 073 (LBG, NAS), R. Long et al. 090806011 (PE), J. Miyake s.n. (NAS), C.M. Zhao EX0291 (PE); Zhuxi, L.Q. Chen IV03548 (CCAU), P.Y. Li 5098 (WUK), 7411 (WUK), 7470 (WUK), 9406 (WUK), 9476 (WUK), 9706 (WUK), 9783 (WUK), 9873 (WUK), 11052 (WUK), 11154 (WUK), S.L. Li GanQL212 (KUN), K.M. Liou 8746 (NAS, WUK); Zigui, C.M. Zhao EX0931 (PE). Hunan: Anhua, K.G. Li et al. AH20140822-2132 (JIU), L. Wu et al. 7561 (CSFI); Baojing, L.H. Liou 9793 (IBSC, NAS); Cili, Q.Y. Liu 4154 (IBSC), W Hunan Exped. 121 (PE), 711 (PE), X.Y. Xi 84226 (PE); Dong’an, X. Li & J. Yang 18110402 (CSFI), J.K. Liu 692 (PE), 810 (PE, P), Y. Liu 826 (HIB, LBG, NAS, PE); Dongkou, Anonymous 614 (IBSC), Z.Y. Yang 245 (IBSC); Fenghuang, Wuling Exped. 1207 (IBSC, PE), Y.G. Yang 3054 (IBSC); Guzhang, L. Su 4331261409070708 (JIU), D.G. Zhang plc002 (JIU); Hongjiang, Anonymous 32 (PE), 425 (IBSC), 3637 (IBSC), Anjiang Agr. Coll. Group 1605 (PE), Z.T. Li 2862 (IBSC, PE); Huayuan, C.B. Ma & L.J. Shi HY20180428_0279 (JIU), HY20180712-0496 (JIU), W.H. Yao 16101636 (CSFI); Jishou, F.L. Teng 16101767 (CSFI), S. Xiang & J.Z. Teng JS20180820-0991 (JIU); Longshan, Y. Xiao et al. LS-216 (CSH), B.M. Yang 2054 (IBSC), D.G. Zhang pph1157 (JIU); Luxi, Q.S. Wang 3698 (HIB), D.G. Zhang 4331221509081047LY (JIU), 4331221510190550LY (JIU), ZZ160815341 (JIU), ZZ170811772 (JIU), ZZ170811773 (JIU); Sangzhi, Anonymous 1223 (KUN), Beijing Exped. 4358 (PE), T.R. Cao 90427 (IBSC), M.X. Jiang et al. 0304 (HIB), W.J. Liu et al. 19092974 (CSFI), Q.S. Wang 2870 (HIB), Z.Z. Yang et al. 0118 (HIB), 0163 (HIB); Shaodong, J.H. Liu 7386 (PE), 7430 (PE); Shimen, Hupingshan Exped. A214 (PE); Suining, K.G. Li & D.G. Zhang SN20150902_ 0376 (JIU), Q.Z. Lin 2002073 (PE); Taoyuan, G.P. Liao & Z.F. Guo 130 (PE), Z.W. Wan 860383 (HUFD); Tongdao, T.C. Chen 653 (IBSC); Wugang, Y.F. Deng 11175 (PE), 11688 (PE), L.H. Liou & G.Z. He 16069 (IBSC, PE, WUK), P.C. Tsoong 1300 (PE); Xinhuang, Wuling Exped. 971 (IBSC, PE); Xinning, K.G. Li xnshs 127 (JIU), xnshs 584 (JIU), L.H. Liou & Q.Z. He 15211 (IBSC, NAS, PE, WUK), L.B. Luo 192 (PE), 1084 (PE), Y.B. Luo 2938 (PE), J.G. Xiao & C.J. Qi 10012 (PE), Q.E. Yang & Y.B. Luo 910038 (PE), Ziyunshan Exped. 90 (PE), 940 (PE), 1131 (PE), 1387 (PE); Xinshao, B.Y. Li 6226 (PE), 6344 (PE); Yongshun, Hunan Exped. 0208 (PE), X.G. Li 204975 (IBK, IBSC), L.H. Liou 9508 (IBSC, NAS, PE), Q.S. Wang 3022 (HIB), W.H. Yao & L.M. Zhu 16101955 (CSFI), D.G. Zhang zdg4331270079 (JIU); Yongsui, Z.S. Ji 19 (NAS); Yuanling, L.H. Liou 08380 (IBSC); Zhangjiajie, X.G. Li 204717 (IBK, IBSC), W.J. Liu et al. 19092636 (CSFI), H. Zhou & D.S. Zhou 15080740 (CSFI), J.J. Zhou 15100305 (CSFI); Zhijiang, Wuling Exped. 1582 (IBSC, PE). Jiangxi: Jiujiang, W. Xiong s.n. (PE); Lushan, S.S. Lai 154 (LBG); Tonggu, S.S. Lai et al. 759 (PE); Wuning, C.M. Tan 8610445 (IBSC), J.H. Zhang 2261 (JJF), 2475 (JJF), 3015 (JJF), J.H. Zhang & Y.Q. Liu TanCM1177 (KUN), J.H. Zhang & D.H. Zhang TCM2837 (JJF), W.Y. Zhao et al. LXP-13-08426 (SYS); Xiushui, JXCM Exped. 36042414029266LY (JXCM), Y.Q. Miao 0505 (SZG), 0711 (SZG), 1188 (JJF), 1902 (JJF), Y.Q. Miao et al. Tancm2124 (JJF), C.M. Tan et al. 14819 (JJF), 081969 (JJF), J. Xiong 07319 (LBG), X.H. Zhan et al. LXP6736 (LBG), S.W. Zhang et al. 360424140710343LY (JXCM). Shaanxi: Baoji, G. Fenzel 79 (WUK); Foping, K.T. Fu 4950 (IBK, PE, WUK); Hanzhong, J.X. Huang & S.M. Xiong HJX10031 (IBSC), Y.L. Qiao 327 (WUK); Langao, Y.S. Chen et al. 1927 (WUK), 2041 (WUK), 3199 (WUK), P.Y. Li 3129 (NAS, WUK), 7662 (SZ, WUK), 7874 (WUK), 7980 (WUK), 8258 (WUK), 8502 (WUK), 8592 (WUK), 10808 (WUK); Lantian, G.X. Su 618 (WUK); Lueyang, K.T. Fu 6042 (IBK, PE, WUK), Z.M. Jiang et al. s.n. (WUK), P.Y. Li 4833 (WUK), S.F. Li et al. 14471 (XBGH), C.L. Tang 286 (WUK), 540 (WUK), 885 (WUK), 975 (WUK), 1151 (WUK), 1171 (WUK); Meixian, Shaanxi Trad. Chin. Med. Exped. 998 (WUK), X.M. Zhang 810 (WUK); Mianxian, K.T. Fu 3890 (WUK), 4021 (WUK), 6021 (PE), P.Y. Li 348 (WUK), 591 (HHBG, WUK); Nanzheng, K.T. Fu 10557 (WUK), 11687 (WUK), Y.C. Ho 32 (WUK), P.Y. Li 432 (WUK), J.W. Wang & Z.C. Shi 145 (PE), T.P. Wang 20763 (WUK); Ningqiang, P.Y. Li 35 (WUK), 110 (WUK), 837 (HHBG, WUK), 4639 (WUK), T.N. Liou & C. Wang 73 (WUK), Y.L. Qiao 63 (WUK), 67 (WUK); Ningshan, K.T. Fu 17803 (IBK, WUK), J.C. Xing 10302 (WUK); Pingli, Y.S. Chen et al. 4204 (WUK), P.Y. Li 4953 (WUK), 8764 (WUK), 9154 (WUK), 9639 (WUK), 10905 (WUK), K.M. Liou 8476 (PE), T.S. Ying 838 (WUK); Shanyang, Anonymous 0240 (HNWP), P.Y. Li 5010 (WUK); Shangnan, S.B. He 628 (WUK), P.C. Kuo 4232 (WUK), 4344 (WUK); Shiquan, J.C. Xing 1069 (WUK), 16302 (WUK); Tongchuan, C.L. Xiao 392 (PE, WUK); Xunyang, P.Y. Li 8939 (WUK), 11002 (WUK); Yangxian, K.T. Fu 5247 (PE, WUK), P.C. Kuo 1844 (IBSC, PE, WUK), 769 (WUK), J.X. Yang 1124 (WUK); Yijun, C.L. Xiao 666 (PE, WUK); Zhashui, X.X. Hou et al. 973 (FJSI); Zhen’an, C.P. Wang 16597 (PE, WUK); Zhenba, Bashan Exped. 4339 (PE), X.S. Jia & J. Yu 3991 (PE); Zhenping, Shaanxi Trad. Chin. Med. Res. Group 2181 (WUK), T.S. Ying et al. 478 (WUK); Zhouzhi, K.M. Liou 8746 (PE); Ziyang, P.Y. Li 4650 (PE, WUK), 4769 (PE, WUK), 4833 (PE, WUK), 4910 (WUK), 4911 (PE), 6944 (WUK), 7024 (WUK), 7215 (WUK), 7263 (WUK). Sichuan: Bazhong, Bazhong Exped. 502 (SM); Baoxing, S.S. Chang & Y.X. Ren 07492 (PE), L.K. Chu 3174 (IBSC, NAS, PE, WUK), K.C. Kuan et al. 2702 (HIB, PE), 2812 (PE), 2879 (PE), 3071 (PE), 3092 (PE), 3111 (PE), Sichuan Agr. Coll. 7492 (IBSC), T.P. Soong 39483 (IBSC, PE, WUK), s.n. (NAS), T.H. Tu 4831 (IBK, IBSC, NAS, PE); Beichuan, C.S. Li & J.X. Huang LCS11325 (IBSC), C.L. Tang et al. 027 (CDBI); Butuo, Anonymous 0736 (SM), 5829 (SM), Sichuan Econ. Plant Exped. 5829 (PE, SM); Cangxi, Sichuan Econ. Plant Exped. 05030 (PE), M.X. Wang 07561 (CDBI, PE); Chengdu, M.Y. Fang s.n. (SZ), Feng et al. 75-hong-1 (FJIDC), H. Li 47112 (HGAS), Xiong et al. s.n. (FJIDC); Chongzhou, Anonymous 0978 (SM), 979 (SM), W.B. Ju et al. AZH00421 (CDBI); Dazhou, Anonymous 0906 (SM); Dechang, Anonymous 0387 (SM); Dujiangyan, C.S. Chang et al. SI0316 (PE), D.Z. Fu et al. 87-1313 (HX), 87-1144 (HX), J.L. Gao 0872450 (HX), D.P. Ho 47112 (IBSC), D.P. Ho & H. Li 47190 (IBSC, PE, SZ), L.C. Hu 21369 (SZ), C.S. Li & J.X. Huang LCS11455 (IBSC), LCS11456 (IBSC), H. Li 46957 (CQNM, IBSC, PE), C. Zhang & G.J. Wu 990245 (HX), D.H. Zhang 1.857 (NAS), D.H. Zhu et al. 952 (PE), 1116 (PE); Ebian, Ebian Exped. 556 (SM), C.T. Kuan 6662 (PE), X.J. Li LiXJ980 (KUN), T.N. Liou 12443 (WUK), Sichuan Econ. Plant Exped. 1505 (CDBI), C.L. Sun 1098 (TIE); Emei, Anonymous 2328 (SM), 2561 (PE), 960643 (PE), W.C. Cheng 6629 (NAS), H.C. Chow 8183 (NAS), Emei Exped. 1691 (SM), W.P. Fang 731 (IBSC), 6009 (IBSC, LBG), 12627 (NAS), 12793 (NAS), 13632 (PE), 17815 (JXU), W.R. He 266 (KUN), Y.Y. Ho 5673 (NAS), 5973 (NAS), C.H. Hsiung et al. 32288 (PE), W.K. Hu 11857 (HGAS), K.C. Kuan et al. 1686 (CDBI, PE, WUK), 2577 (PE), 2625 (PE), 2739 (PE), 04164 (CDBI, PE, WUK), T.C. Lee 3548 (TIE), T.N. Liou 12443 (WUK), T.N. Liou & C. Wang 823 (WUK), Y.S. Liu 1074 (LBG), Sichuan Univ. Biol. Coll. Exped. 52174 (PE), Y.H. Tau 51447 (AU, IBSC), 51683 (AU, IBSC), 51722 (AU, IBSC), J.Q. Tian & J.J. Liu 0084 (PE), 0181 (PE), H.L. Tsiang & S.S. Chang 32288 (IBK), T.H. Tu 698 (IBK, IBSC, NAS, PE, WUK), 3704 (IBSC), Z.R. Wang & Z.X. Zhang C0153 (PE), H.G. Xu 2801 (PE), K.H. Yang 56861 (IBSC, NAS, PE), 57231 (IBSC, NAS, PE), C.W. Yao 2879 (NAS), 3385 (NAS), 5224 (IBK, NAS), G. Zhang 1160 (NAS), X.B. Zhang 293 (NAS), 32288 (LBG), Z.X. Zhao 727 (NAS), G.S. Zhou 81212 (PE), Z.Y. Zhu 275 (WUK); Ganluo, Anonymous 875 (SM), 4217 (SM), Sichuan Econ. Plant Exped. 4217 (PE), 4437 (PE); Gaoxian, Anonymous 492 (SM); Gongxian, Anonymous 533 (SM); Gulin, J.H. Chen et al. 4449 (SM), 4497 (SM), Gulin Group 653 (SM), C.S. Li & J.X. Huang LCS11084 (IBSC), LCS11152 (IBSC), PE-Gulin Exped. 0482 (PE), 0702 (PE); Guangyuan, C.Y. De 4184 (CDBI), Guangyuan Exped. 0596 (SM), W.K. Hu 52 (SZ), K.C. Kuan et al. 3156 (PE), 3157 (PE); Hanyuan, H.C. Chow 535 (HIB), Hanyuan 0783 (SM), T.P. Wang 8827 (IBSC, PE), Z.C. Zhao 0413 (PE), 0554 (PE); Hechuan, Hechuan Exped. 0106 (SM), T.H. Tu 5193 (IBSC, PE); Hejiang, J.H. Chen et al. 4775 (SM), Hejiang Group 0515 (SM), 0575 (SM); Hongya, W.P. Fang 8389 (LBG, NAS, PE), 24617 (NAS), Hongya Group 1226 (SM), C.W. Yao 2706 (NAS), 2829 (NAS); Huaying, C. Pei 7149 (NAS); Huidong, Huidong Exped. 0465 (SM), C.S. Yok et al. 4187 (NAS), 64187 (NAS); Huili, W.P. Fang 9524 (IBSC, NAS, PE), C. Ho et al. 11527 (NAS, SM), 11667 (NAS), R.G. Li 0400 (SM), T.T. Yu 1366 (NAS, PE), 1610 (PE); Jiange, Hechuan 526 (SM); Jianyang, Jianyang Group 2620 (SM), Y.P. Hsu 462 (HHBG, WUK); Jinyang, Anonymous 038 (SM); Jiulong, Anonymous 0298 (SM), Jiulong Exped. 0302 (SM), 0391 (SM), G.F. Li 63122 (IBSC, NAS), Q.Q. Wang 21424 (CDBI); Jiuzhaigou, Anonymous 1149 (SM); Junlian, Anonymous 1288 (SM); Kangding, W.K. Hu & C. Ho 11468 (PE); Leshan, L.Y. Tai 163 (NAS); Lezhi, Anonymous 1559 (SM); Leibo, Anonymous 614 (SZ), 27888 (SM), C.T. Kuan 8314 (PE), Leibo Trad. Chin. Med. Exped. 0994 (SM), C.S. Li & J.X. Huang LCS11184 (IBSC), Q.S. Zhao & M.Y. He 118631 (CQNM), 122892 (CDBI, CQNM), Q.S. Zhao & J.B. Shi 119433 (CDBI, CQNM), Q.S. Zhao & C.M. Tan 118068 (CDBI, CQNM), 119089 (CDBI, CQNM), 123436 (CDBI, CQNM), 123568 (CDBI, CQNM); Lixian, C. Ho & T.L. Chow 13888 (IBSC, LBG, NAS, PE); Longchang, Longchang Exped. 92 (SM); Luding, Anonymous 15906 (SM), D.E. Boufford et al. 27409 (PE), W.K. Hu & C. Ho 11468 (HGAS, WUK), K.C. Kuan et al. 1666 (HIB, PE), Sichuan Trad. Med. Res. Exped. 15906 (NAS, SM), Sichuan Veg. Exped. 41586 (CDBI), C.Y. Wu et al. 6259 (KUN), G.H. Xu 25792 (CDBI), 26266 (CDBI), Veg. Group 31930 (CDBI), 41281 (CDBI), 41323 (CDBI), 41811 (CDBI), 46075 (CDBI), 46549 (CDBI); Lushan, Lushan Exped. 1003 (SM), D.Y. Peng 46814 (IBSC), 47959 (CDBI); Luxian, W.P. Fang 9818 (NAS, PE); Luzhou, Z.C. Zhong 6054 (CDBI); Mabian, Mabian Group 1168 (SM), 1169 (SM), S.L. Sun 5861 (PE), T.T. Yu 5861 (IBSC, PE); Maoxian, S.Y. Chen et al. 5916 (SM), W.P. Fang 5533 (NAS), 5535 (LBG), X.F. Gao & Y.L. Peng 1548 (CDBI), 1922 (CDBI), C. Ho 5916 (NAS), C. Ho & T.L. Chow 13888 (IBSC, LBG, NAS, PE), C.S. Li & J.X. Huang LCS11309 (IBSC), Maoxian-Wenchuan Group 5151 (CDBI, SM), Sichuan Veg. Exped. 8732 (CDBI), W.P. Wang 5533 (IBSC, LBG, NAS, PE), T.T. Yu 2504 (NAS, PE); Meigu, Anonymous 1728 (SM), Sichuan Econ. Plant Exped. 1091 (PE), 1364 (PE), 1728 (KUN, PE), C.T. Kuan 9008 (PE), Meigu Exped. 0512 (SM), Sichuan Veg. Group 13025 (CDBI, PE); Mianning, W.P. Fang 9497 (IBSC, NAS, PE), Mianning Exped. 0475 (SM), S.F. Zhu 20495 (PE); Mianyang, Y.P. Hsu 49 (HHBG, WUK), 213 (WUK), Mianyang Exped. 214 (SM); Miyi, Anonymous 306 (SM), S.Y. Chen et al. 11009 (NAS, SM); Muchuan, Anonymous 0806 (SM), Muchuan Group 0126 (SM); Muli, S.Y. Chen et al. 11009 (SM), G. Hu & F.Q. Lü 7787 (CDBI, CDCM), J.S. Liou 7672 (CDBI, CDCM), Qinghai-Xizang Exped. 13308 (PE), T.T. Yu 14042 (PE), 14728 (PE), Q.S. Zhao 8638 (CDBI, CDCM), Q.S. Zhao et al. 7519 (CDBI), 7565 (CDBI); Nanchong, Sichuan Econ. Plant Exped. 00034 (CDBI, PE, SM), 00045 (PE, SM); Nanjiang, Anonymous 820 (SM), Sichuan Econ. Plant Exped. 00214 (CDBI); Naxi, Q. Yu et al. TXD-D-0122 (CDBI); Neijiang, Y.P. Hsu 386 (HHBG, WUK); Ningnan, Anonymous 0510 (SM); Pengxi, Anonymous 515 (SM); Pingchang, Anonymous 672 (SM); Pingshan, Anonymous 1059 (SM), Sichuan Econ. Plant Exped. 0765 (CDBI, PE); Pingwu, W.P. Fang 4438 (NAS, PE), C.S. Li & J.X. Huang LCS11337 (IBSC), LCS11342 (IBSC), Pingwu Exped. 525 (SM), H.L. Tsiang 11112 (NAS, PE), S.Z. Zhang et al. 929 (PE); Puge, Anonymous 5542 (SM), 25212 (SM), Puge Exped. 484 (SM), 839 (SM), Sichuan Econ. Plant Exped. 5542 (PE), Sichuan Trad. Med. Res. Exped. 25212 (NAS, SM), 25862 (NAS); Qingchuan, Anonymous 2612-2 (SM), 2612-3 (SM); Qingshen, Qingshen Exped. 547 (SM); Qionglai, Anonymous 1249 (SM); Quxian, Anonymous 0495 (SM), Z.Y. Liu 4646 (SM); Renshou, Renshou Group 0529 (SM); Santai, Santai Group 404 (SM); Shehong, M. Yuan YUANM2015L065 (KUN); Shifang, Anonymous 0881 (SM), 0886 (SM), 0903 (SM), 0904 (SM); Shimian, C.C. Hsieh 41484 (IBSC), 42366 (IBSC, PE), 42382 (IBSC, PE), 42484 (PE, SZ), 42678 (IBSC, PE), H.S. Kung 42382 (WUK), 42483 (WUK), Shimian Exped. 1109 (SM), J. Wang & C. Luo LuoP210 (KUN), G.H. Xu 25649 (CDBI), Z.C. Zhao 1320 (PE), 1524 (PE), 1655 (PE), 1767 (PE), 1952 (PE), 1970 (PE), 2073 (PE), 2351 (PE), 2355 (PE), 2677 (PE), 2832 (PE), 2915 (PE), 3079 (PE), 3273 (PE); Suining, X.H. Hu 141 (CDBI, PE), Suining Group 365 (SM); Tianquan, K.L. Chu 4133 (IBSC, NAS, PE), Y.Y. Ho 6854 (NAS), W.K. Hu & C. Ho 11857 (PE, SZ), K.C. Kuan et al. 2042 (PE), 2395 (HIB, PE), D.Y. Peng 46691 (IBSC), 46907 (IBSC), F.C. Tai & C.M. Teng 5083 (SZ), Tianquan Exped. 759 (SM), 805 (SM), H.L. Tsiang 35228 (CQNM, IBK, IBSC, PE), 35302 (IBK, IBSC, PE), 37361 (IBK), T.H. Tu 4598 (IBSC, NAS, PE); Tongjiang, Tongjiang Exped. 749 (SM), J.A. Wang 0172 (CDBI); Wanyuan, Anonymous 1152 (SM), Bashan Exped. 3672 (PE), C.Y. Chang 0116 (CDBI), X.S. Jia & J. Yu 3931 (PE), P.Y. Li 4225 (PE, WUK), 4426 (PE, WUK), 4639 (PE), 5690 (WUK), 5806 (PE), 5940 (WUK), 5982 (WUK), 6040 (WUK), Wanyuan Group 0600 (SM); Weiyuan, Z.W. Wu 630 (SM); Wenchuan, H.C. Chow 1071 (NAS), K.Y. Lang et al. 1246 (PE), 1372 (PE), Veg. Group 30670 (CDBI), C. Zhang et al. 20060398 (HX); Xichang, W.P. Chen 0430 (SM), T.T. Yu 1231 (IBSC, NAS, PE); Xide, Anonymous 0567 (SM); Xingwen, Xingwen Exped. 363 (SM); Xuyong, J.H. Chen et al. 4423 (SM), W.B. Ju & H.N. Deng HGX12608 (CDBI), HGX12684 (CDBI), HGX13488 (CDBI), HGX13555 (CDBI), S.G. Tang 1453 (SM), Xuyong Exped. 1022 (SM), K.P. Yin & Q.H. Chen 0632 (CDBI); Ya’an, W.P. Fang 8940 (LBG, NAS), Y.Q. Hao BX-60 (CSH), Mingshan Exped. 0672 (SM), Ya’an Exped. 0814 (SM); Yanbian, Yanbian Exped. 0368 (SM); Yanting, Anonymous 6073 (CDBI); Yibin, Yibin Exped. 1137 (SM); Yingjing, Yingjing Exped. 0482 (SM); Yuexi, Anonymous 0772 (SM), 26112 (SM), 26530 (SM), W.P. Fang 4989 (NAS), C.S. Li & J.X. Huang LCS11213 (IBSC), LCS11245 (IBSC), Sichuan Econ. Plant Exped. 3495 (PE), 4534 (PE), Sichuan Trad. Med. Res. Exped. 26012 (NAS), 26112 (NAS, SM), 26530 (NAS, SM), 27888 (NAS, SM); Zhaohua, Y.C. Ho 1780 (WUK), Z.P. Wei 3826 (WUK); Zhaojue, Sichuan Econ. Plant Exped. 128 (CDBI), Zhaojue Exped. 0879 (SM); Zhongjiang, Zhongjiang Exped. 327 (SM); Ziyang, Y.P. Hsu 419 (HHBG, WUK), 462 (HHBG, WUK). Taiwan: Chiayi, Anonymous s.n. (TAI), T.C. Huang et al. 14308 (TAI), T. Nawada s.n. (TAI), T. Suzuki 19364 (PH), H. Takasu s.n. (TAI); Hsinchu, S.Y. Huang et al. 660 (TAI), Y.F. Wang 698 (TAI); Hualien, S.F. Huang et al. 5189 (TAI), S.F. Huang et al. 226718 (TAI), S. Sasaki s.n. (TAI); Ilan, H.Y. Chen et al. 186 (TAI), S.F. Huang 4782 (TAI), S. Suzuki 4105 (TAI), s.n. (TAI), H.T. Yu 2068 (P); Kaohsiung, S. Sasaki s.n. (TAI); Miaoli, R.J. Ou et al. 2213 (TAI), C.H. Yu 818 (TAI); Nantou, S.T. Chiu & K. C. Yang 04612 (PE), H.N. Qin 18799 (PE), S. Sasaki s.n. (PH, TAI), C.M. Wang et al. 03309 (PE); Pingtung, S. Sasaki 39088 (PH); Taitung, Y. Yamamoto & K. Mori Y.M.537 (PH); Taoyuan, S.Y. Huang s.n. (TAI), T.Y.A. Yang et al. 06251 (PE). Yunnan: Binchuan, M. Chen 2316 (NAS), R.C. Ching 24915 (PE); Daguan, NE Yunnan Exped. 484 (PE); Dali, M. Chen 2627 (NAS), X.F. Gao et al. 36 (KUN), 105 (KUN), 137 (KUN), T.N. Liou 14582 (IBSC, KUN), 14893 (IBSC, KUN, PE), 17493 (NAS), 21449 (IBSC, PE), X.L. Hou et al. 331 (AU), Z.Y. Su et al. 1145 (KUN), H.C. Wang 1205 (IBSC, KUN, PE), Q.E. Yang 9412 (PE); Deqên, Z.D. Fang et al. SN2007-16 (SABG), K.M. Feng 5550 (PE), 5955 (PE), IBCAS Hengduanshan Exped. 3717 (PE), Kunming Work. Group 23577 (PE), T.T. Yu 9641 (PE), 10283 (PE), 10419 (IBSC, PE), 10552 (PE), 10577 (PE), C.W. Wang 69127 (IBSC, NAS, PE), 69399 (IBSC, PE); Dongchuan, NE Yunnan Exped. 472 (KUN); Eryuan, P.Y. Kiu 60987 (CDBI, KUN), 61017 (CDBI); Fugong, Gaoligong Shan Biodiversity Sur. 25581 (PE), 27681 (PE), H.T. Tsai 54661 (NAS), 56629 (NAS), 58831 (NAS, PE), 59079 (LBG, NAS, PE); Fumin, P.Y. Kiu 596018 (PE), 771289 (IBK); Funing, Funing Exped. 5326281606221444LY (IMDY); Guangnan, J.X. Wang & C.T. Ting 1027 (NAS); Gongshan, Gaoligong Shan Biodiversity Sur. 23222 (PE), C.W. Wang 66664 (LBG, NAS, PE, WUK), T.T. Yu 23056 (PE), 23155 (PE); Heqing, R.C. Ching 23807 (PE), 24078 (PE); Jingdong, P.Y. Kiu 53471 (KUN), Z.H. Yang et al. 101449 (KUN); Kunming, Anonymous 1039 (PE), 1079 (PE), 2128 (PE), P.Y. Kiu 51392 (IBSC), 54950 (IBSC, PE), 55051 (PE), 77714 (CDBI), Kunming Work. Group 51392 (PE), S.B. Lan 253 (PE), G.X. Liang 283 (IBSC), W.S. Liou 51392 (IBSC), P.I. Mao 77 (KUN, PE), S.K. Wu 38 (IBK); Lanping, H.T. Tsai 53793 (PE, WUK), 56027 (PE, WUK), 57863 (NAS), 58027 (NAS); Lijiang, R.C. Ching 21261 (PE), K.M. Feng 21677 (PE), G. Forrest 2936 (PE), Guo 2009 (KUN), H.C. He 21859 (PE), IBCAS Hengduanshan Exped. 02685 (PE), Kunming Work. Group 22653 (PE), G.S. Qin 309 (XBGH), J.F. Rock 6137 (P), K. Wu & J.C. Hao YN211 (BNU), Z.H. Yang 84128 (IBSC), T.T. Yu 15587 (PE), Yulong Exped. 5307210263 (IMDY), Yunnan Univ. Biol. Coll. 00814 (IBSC), Y.Z. Zhao 22013 (NAS); Luquan, P.I. Mao 1562 (WUK); Mengzi, C.W. Wang 83310 (KUN); Pingbian, H.T. Tsai 61782 (PE), 62597 (P), C.W. Wang 82117 (PE); Qiaojia, T.B. Zhang SCSB-W-833 (KUN); Shangri-la, Z.D. Fang et al. K-1895 (SABG), K.M. Feng 2053 (PE), Q.E. Yang & Q. Yuan Yangqe2327 (PE), T.T. Yu 10942 (PE), Zhongdian Exped. 1492 (PE); Shilin, Y.M. Shui et al. 65169 (PE); Tengchong, J. 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Notes:— Anemone robustostylosa , having purple-red flowers, is further referable to A. hupehensis f. hupehensis , while A. discoideogyna , having white flowers, is further referable to A. hupehensis f. alba Wang (1974: 166) . The flowers in A. hupehensis , however, vary in color in a continuous manner within and between populations, with different intermediates existing between the purple and white ones ( Figs. 4 View FIGURE 4 , 8 View FIGURE 8 , 9 View FIGURE 9 ). The recognition of f. alba and f. hupehensis under A. hupehensis seems not to be significant biologically.

As elaborated on by Bowles & Stearn (1947), the name A. hupehensis was applied originally to a plant of Chinese origin offered to the public in the autumn of 1908 by the well-known French nursery V. Lemoine & Fils of Nancy as A. japonica hupehensis , with only a descriptive note in French, and in 1910, the Lemonines treated A. japonica hupehensis as an independent species of its own and named it A. hupehensis , also with only a descriptive note (it is noteworthy that A. japonica hupehensis was actually not explicitly ranked as a variety then ). Although it is clear that taxonomically A. hupehensis , as the name indicates, was first known from Hubei province in central China, we have been unable to trace any original material for this name. This species is indeed very common in Hubei. It is fortunate that the identity of A. hupehensis has long been well established and there is no problem with the correct application of the name A. hupehensis . We do not think it quite necessary and urgent to designate a neotype for this name herein.

Of the specimens cited as Anemone hupehensis by Wang (2021), the collection Z.X. Tang 1507 ( Fig. 13A–C View FIGURE 13 ) from Maowen (now Maoxian and Wenchuan) in western Sichuan, China, actually belongs to A. tomentosa ( Maximowicz 1889: 7) Pei (1933: 2) . These two species are most closely similar to each other, with the former different from the latter by having leaves abaxially strigose (vs. tomentose). Moreover, a collection from Wenshan in southeastern Yunnan, China, Y.M. Shui 3365 (PE; Fig. 13D View FIGURE 13 ), was wrongly cited as Y.M. Shui 336.