Plusioglyphiulus, SILVESTRI, 1923

KEY TO SPECIES OF PLUSIOGLYPHIULUS SILVESTRI, 1923 View in CoL

1. At least some crests on collum complete ( Figs 4A, B; 21A, B) .................................... 2

— No complete crests on collum, all interrupted ( Figs 1A, B; 8A, B; 14A) ..................... 5

2. Collum with 3+3 complete crests dorsad of lateralmost crests, which are frontally abbreviated; carinotaxy pattern mixostichic. Dorsal tubercles of second row on midbody metaterga displaced, intercalary, not in line with tubercles of the other two rows. Sabah................................................................................................................ P. macfarlanei View in CoL

— Collum with a different number of complete crests dorsad of lateralmost crests; carinotaxy pattern isostichic (i.e. devoid of intercalary tubercles or crests) ( Figs 4A, B; 21A, B). Dorsal tubercles or crests arranged in clear-cut longitudinal rows ( Fig. 21C, D) .................... 3

3. Body mostly pigmented, usually brown with a pattern of yellowish markings; head between ocellaria at least partly brown. Midbody metaterga with 3/3/3+m/m+3/3/3 tubercles between poriferous cones. Male telopodites 3 stout and compact. Posterior gonopods with processes ............................................................................................................. 4

— Body completely unpigmented. Carinotaxy formula of midbody segments 3/3+I/i+2/2+m/m. Male telopodites 3 not compact, rather elongated. Posterior gonopods without processes. Cave in Sarawak..................................................................................... P. cavernicolus View in CoL

4. Body length 30-35 mm, width 2.0- 2.2 mm. Collum with 3+3 complete crests, including lateralmost ( Fig. 4A, B). Carinotaxy formula of midbody metaterga 3/3/3+I/i+3/3/3+m/m ( Figs 4D; 5B). Male legs 3 strongly enlarged, typical ( Figs 5D; 7D). Gonopods complex ( Figs 6; 7G, H). South Vietnam...................................................... P. ampullifer View in CoL n. sp.

— Body length about 22 mm, width 1.3 mm. Collum with 5+5 complete crests in addition to lateralmost ( Fig. 21A, B). Carinotaxy formula of midbody metaterga 4/4+I/i+3/3/3+m/m ( Figs 21C, D; 22A). Male legs 3 less enlarged ( Fig. 23C). Gonopods simple ( Fig. 23 F-H). Kalimantan.......................................................................................... P. bedosae View in CoL n. sp.

5. Carinotaxy pattern of collum isostichic (i.e. devoid of intercalary tubercles or crests). Epiproct with 1+1 high subtransverse tubercles and a median knob ( Fig. 14E, F). Gonopods highly complex ( Figs 16; 17F, G). Laos................................................ P. steineri View in CoL n. sp.

— Carinotaxy pattern of collum mixostichic. Epiproct without high subtransverse tubercles, a median knob usually missing, paramedian crests or tubercles longitudinal. Gonopods simpler ....................................................................................................................... 6

6. Carinotaxy formula of midbody metaterga 3/3+I/i+3/3+m/m ( Fig. 1C, D). Anterior gonopods with a prominent, spiniform, median, sternal process (st) ( Fig. 3C, D). Caves in Malaya ................................................................................................ P. grandicollis View in CoL

— Carinotaxy formulae of midbody metaterga 3/3+I/i+3/3(4)/3+m/(m)/m. Anterior gonopods without sternal process ............................................................................................... 7

7. Second transverse row of tubercles on midbody metaterga intercalary, apparently developed only towards onset of sexual maturity, formula 3/3+I/i+3/4/3+m//m. Epiproct short ( Figs 27F; 32A). Legs more or less as long as body diameter. Borneo .......................... 8

— Dorsal tubercles or crests arranged in clear-cut longitudinal rows, pattern isostichic. Epiproct longer. Legs usually closer to being half as long as body diameter ............... 10

8. Body length 43-50 mm, width 2.0- 2.1 mm; segments 64-70p+2-1a+T. Collum with a distinct transverse impression near anterior third. Carinotaxy formula of midbody segments 2/2+I/i+3/4/3+m//m (NB: Fig. 24C, D shows juvenile condition). Paraprocts flattened. Kalimantan........................................................................................ P. pallidior View in CoL n. sp.

— Body length up to about 30 mm, width 1.0- 1.6 mm; segments up to 53p+1a+T. Collum not particularly impressed near anterior third. Carinotaxy formula of midbody segments 3/3+I/i+3/4(5)/3+m//m. Paraprocts regularly convex ................................................. 9

9. Brown spots present dorsad of ocellaria. Epiproct very short, roundly subtrapeziform ( Figs 31E, F; 32A). Anterior gonopods without cxp2 ( Fig. 33C). Arms (d) of posterior gonopods rudimentary ( Fig. 33D). Sarawak...................................... P. hoffmani View in CoL n. sp.

— Brown spots absent dorsad of ocellaria. Epiproct a little longer, regularly rounded ( Fig. 27E, F). Anterior gonopods swollen, stout, with both cxp1 and cxp2 ( Figs 29B, C; 30G, H). Arms (d) of posterior gonopods long, spiniform and lateral in position ( Figs 29D; 30I). Kalimantan................................................................................... P. similis View in CoL n. sp.

10. General coloration very dark brown to blackish (fading to reddish after long conservation in alcohol). Paraprocts with a distinct, median, ridge-like elevation. Cambodia... P. dubius View in CoL

— General coloration light brown to brown. Paraprocts flat medially ........................... 11

11. Paraprocts flattened ( Fig. 19B). Posterior gonopods with a rather long, spiniform arm (d) ( Fig. 20G) ....................................................................................... P. deharvengi View in CoL n. sp.

— Paraprocts regularly convex. Posterior gonopods without spiniform arm .................. 12

12. Epiproct broadly and regularly rounded ( Figs 8E, F; 9A). Posterior gonopods with rudimentary arms (d) ( Fig. 10G) ......................................................... P. foveatus View in CoL n. sp.

— Epiproct roundly subtrapeziform. Posterior gonopods different ................................ 13

13. Brown spots present dorsad of ocellaria. Collum strongly impressed transversely in anterior third. Male legs 1 with very small coxites. Anterior gonopods with very short and lobe-like cxp1. Posterior gonopods without arms. Cambodia....................................... P. boutini View in CoL

— Brown spots absent dorsad of ocellaria. Collum not particularly depressed near anterior third. Male legs 1 with large coxites ( Fig. 13B). Anterior gonopods with very long, acuminate cxp1 ( Fig. 13F). Posterior gonopods with short, sacciform arms (d) ( Fig. 13G). Thailand.............................................................................................................. P. bessoni View in CoL n. sp.