Plusioglyphiulus deharvengi, Golovatch & Geoffroy & Mauriès & Den Spiegel, 2009

Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul & Den Spiegel, Didier Van, 2009, Review of the millipede genusPlusioglyphiulusSilvestri, 1923, with descriptions of new species from southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae), Zoosystema 31 (1), pp. 71-116 : 93-97

publication ID

https://doi.org/10.5252/z2009n1a5

DOI

https://doi.org/10.5281/zenodo.17940871

persistent identifier

https://treatment.plazi.org/id/03D71761-B662-FFF6-FF74-F9E7127AFB44

treatment provided by

Marcus

scientific name

Plusioglyphiulus deharvengi
status

sp. nov.

Plusioglyphiulus deharvengi View in CoL n. sp. ( Figs 18-20)

TYPE MATERIAL. — Laos. Luang Prabang Prov., 19.88°N, 102.14°E, Xieng Maen, Cave Vat Tham , by hand, 30.XII.1999, leg. L. Deharveng & A. Bedos (LAO-106), holotype ♂ ( MNHN GA059) GoogleMaps ; 1 ♂, 1 ♀ paratypes ( MNHN GA059); 1 ♀ paratype ( ZMUM); 1 ♀ paratype ( SEM).

ETYMOLOGY. — To honour Louis Deharveng, one of the collectors and donators of this material.

DIAGNOSIS. — Differs from congeners by certain details of gonopod structure (complex anterior gonopods versus very simple posterior ones), coupled with the mixostichic carinotaxy of the collum, the isostichic carinotaxy of other body segments, the flattened paraprocts, the elongated and subtrapeziform epiproct and the presence of spots dorsad of the ocellaria.

DESCRIPTION

Length of adults 20-33mm,width 1.3-1.9mm,collum and segments of posterior third of body being equally broadest.Coloration usually uniformly yellow-brown, sometimes with a paramedian pair of more or less wide,grey-yellowish stripes;collum,head and segment 2 mostly pale; tips of tergal crests/tubercles usually dark brown; ocellaria brown to blackish, with a pair of adjacent,light brown,well-separated spots dorsally; antennae and distal podomeres light brownish.

Body with 48-59p+4-2a+T. Length of holotype about 33 mm, width 1.9 mm, with 59p+2a+T.

Antennae short and clavate ( Fig. 20A), antennomeres 6 and 7 each with a modest group of bacilliform sensilla distodorsally. Ocellaria transversely subreniform, with 8-11 ocelli in 5-7 longitudinal rows. Gnathochilarium oligotrichous, each lamella lingualis with three setae distally.

Postcollar constriction evident, but collum and segment 2 not particularly strongly enlarged ( Fig.18B). Carinotaxy formula of collum slightly variable: ta/ t+/(t)/+2p/t+(ta)/(t)+3p/t+4p+ta/t/t/t+6p/t/t+ta/ t+pp/t/t+ma/t (usually), ta/t+2p/t+3p/t/t+ta/t+5p+ta/ t+7p/t/t+ta/t+pp/t/t+ma/t or ta/t+2p/t+(ta)/(t)+3p/ t+(ta)/(t)+4p+ta/t/t/t+6p/t/t+ta/t+pp/t/t+ma/t (more rarely) ( Fig. 18A, B). Carinotaxy of metatergum 2, 8/8+m/m+8/8; formula of metaterga 3 and 4, 7/7+m/m+7/7; formula of metaterga 5 to 16 and of several segments before apodous, 3/3+I/i+3/3/3+m/ m+3/3/3+I/i+3/3; formula of midbody metaterga, 3/3+I/i+3/3/3+m/m/m+3/3/3+I/i+3/3; that of apodous segments, 4+3/3+m/m+3/3+4 ( Figs 18; 19A); all crests and tubercles rather low, poriferous cones (I) considerably higher, about as high as broad, while median ones (m) lower than others ( Figs 18; 19C). Dorsal crests on several posteriormost segments considerably higher than others ( Fig. 19A). Midbody segments circular in cross-section ( Fig. 19C). Pleural regions of segments 2-4 conspicuously expanded, flap-shaped, especially so on segment 3 ( Fig. 18A). Pleural ridge behind gonopod aperture on male segment 7, tegument and limbus as in above congeners. Epiproct elongate and subtrapeziform apically, with a paramedian pair of high longitudinal crests/ tubercles flanked on each side by 1 or 2 small knobs at about midway; paraprocts flattened; hypoproct emarginate at caudal margin ( Fig. 19A, B).

Legs short and rather stout, on midbody segments about half as long as body diameter ( Figs 19C; 20E). Claw at base with a strong but relatively short accessory spine about ¼ as long as claw itself ( Fig. 19D). Tarsi, their distal setae and male legs 1-3 ( Fig. 20 B-D) as in P. grandicollis .

Anterior gonopods ( Fig. 20F) much like in P. ampullifer n. sp., but processes cxp1 non-contiguous, lamelliform, broad and high, with a few setae laterally near base, subsecuriform, terminating in a minute uncus apico-medially and a long, digitiform, slightly curved outgrowth apico-laterally; processes cxp2 of similar shape and length, but retrorse at end; telopodites (te) simple, laterocaudal in position, movable, rather slender and high, subcylindrical, setose in distal half laterally. Posterior gonopods ( Fig. 20G) small, simple, without traces of telopodites; each coxite with a rather long, spiniform arm (d) at about midway on frontal face, distal half lamelliform, parallel-sided, with a deep fovea and a modest emargination apically.

REMARKS

Any obvious troglomorphic traits in this species seem to be absent.

MNHN

Museum National d'Histoire Naturelle

ZMUM

Zoological Museum, University of Amoy

Kingdom

Animalia

Phylum

Arthropoda

Class

Diplopoda

Order

Spirostreptida

SubOrder

Cambalidea

Family

Cambalopsidae

SubFamily

Glyphiulinae

Genus

Plusioglyphiulus

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