Nitidiscala detracta

Nitidiscala detracta (De Boury in Cossmann, 1912)

Figs 33L View FIGURE 33 , 37E–H View FIGURE 37

Scalaria (Hirtoscala) muricata (Risso) View in CoL — Boettger 1902: 85 [non Scalaria muricata Risso, 1826 View in CoL ].

* Clathrus detractus De Boury nom. mut.— Cossmann 1912: 172, pl. 5, fig. 4.

Scala (Clathrus) detracta De Boury— Zilch 1934: 231 , pl. 10, fig. 81.

Scala (Clathrus) detracta De Boury — Csepreghy-Meznerics 1956: 390, pl. 3, figs 46–47.

non Scala (Clathrus) detracta (De Boury in Cossmann, 1912)— Bałuk 1975: 172, pl. 20, fig. 14 [= Hirtoscala cornucopiae sp.

nov.].

Type material. Holotype, SMF 380172_12-1448a ( XII 1448 a), illustrated in Zilch (1934: pl. 10, fig. 80), SL: 6.8 mm, MD: 3.5 mm, CoŞteiu de Sus ( Romania), illustrated in Zilch (1934: pl. 10, fig. 80), Fig. 37E View FIGURE 37 .

Illustrated material. NHMW 1865/0035/0082, SL: 5.1 mm, MD: 3.2 mm, Szob ( Hungary), Figs 37G View FIGURE 37 1 –G View FIGURE 1 2 View FIGURE 2 . NHMW 1997z0178/0973, SL: 4.8 mm, MD: 2.8 mm, Bad Vöslau ( Austria), Figs 33L View FIGURE 33 , 37F View FIGURE 37 1 –F View FIGURE 1 2 View FIGURE 2 . NHMW 1997z0178/ 0973a, SL: 4.2 mm, MD: 2.6 mm, Bad Vöslau ( Austria), Fig. 37H View FIGURE 37 .

Revised description. Shell small, moderately broad (apical angle ~31°), weakly coronate. Protoconch and early teleoconch whorls unknown. Teleoconch of at least seven strongly convex whorls with periphery placed just above mid-whorl. Suture very deeply impressed. Sculpture of widely spaced, strongly raised, thin, prosocline lamellae, obliquely aligned axially (7: 9: 8), lamellae forming short, vaguely trigonal spines along shoulder. Crests of lamellae erect. Interspaces wide, smooth, glossy. Last whorl slightly widening, attaining about 40% of total height, with shallow subsutural ramp, strongly convex below, lamellae strongly rolled over on base, persisting to columellar callus, no peribasal cord but lamellae form angled nodes at transition to convex base. Very short, broad fasciole.Aperture subcircular, columella weakly convex. Outer lip thickened by varix. Columellar and parietal callus forming broad rim, detached from base and fasciole. Umbilicus narrow, open on subadult specimens (holotype), fully covered by columellar callus on adult specimens.

Discussion. This species was placed in Nobiliscala by de Boury (1917: 60). It was confused by Boettger (1902) with the extant western Atlantic Hirtoscala foliaceicosta (d’Orbigny, 1842) (as Scalaria muricata G. B. Sowerby II, 1844 ) (see Weil et al. 1999: fig. 35). Both species are indeed very similar, but the lamellae of H. foliaceicosta are erect over the base and not rolled over as in Nitidiscala detracta (De Boury in Cossmann, 1912). A specimen illustrated as this species by Csepreghy-Meznerics (1956) from Szob ( Hungary) is slenderer and has a narrower subsutural ramp. We have not seen this specimen and include it here with some hesitation. However, another specimen of N. detracta from Szob is illustrated herein, proving its occurrence at that locality. The specimen illustrated by ( Bałuk 1975) from Korytnica ( Poland) differs in its much broader subsutural ramp and prominent shoulder spines. That specimen is identified herein as Hirtoscala cornucopiae sp. nov.

Paleoenvironment. Unknown.

Distribution in Central Paratethys. Badenian (Middle Miocene): Vienna Basin: Bad Vöslau ( Austria) (hoc opus); Pannonian Basin: Szob ( Hungary) (hoc opus); Făget Basin: CoŞteiu de Sus ( Romania) ( Boettger 1902).