Anguloacrilla lima, Harzhauser & Landau, 2025

Anguloacrilla lima sp. nov.

Figs 7A View FIGURE 7 , 13B View FIGURE 13 1 –B View FIGURE 1 2 View FIGURE 2

Type material. Holotype, NHMW 1865 View Materials /0001/0244, SL: 28.6 mm, MD: 11.3 mm, Lăpugiu de Sus ( Romania), Figs 7A View FIGURE 7 , 13B View FIGURE 13 1 –B View FIGURE 1 2 View FIGURE 2 .

Type locality. Lăpugiu de Sus ( Romania), Făget Basin .

Type stratum. Silt and clay of the Dej Formation.

Age. Middle Miocene, Badenian (Langhian).

Etymology. After lima (Latin for file), referring to the sculpture; noun in apposition.

Diagnosis. Large, moderately slender shell with low, convex whorls, very dense sculpture of granulose lamellae and numerous delicate spiral threads in narrow axial interspaces, broad, well-developed abapical apertural spout.

Description. Shell large, moderately slender (apical angle ~27°). Protoconch and early teleoconch whorls unknown. Teleoconch of at least eight relatively low whorls. Early spire whorls convex with distinctly impressed suture. Later whorls moderately convex with very narrow subsutural ramp. Periphery placed mid-whorl. Sculpture of numerous, thin, low, densely spaced, weakly prosocline lamellae, not aligned axially (33: ~100: ~90). Crests of lamellae erect. Interspaces between lamellae with numerous, prominent spiral threads, separated by interspaces of equal width, resulting in extremely fine cancellate pattern. Last whorl attaining 35% of total height, marked basal angulation delimited by narrow peribasal cord, flattened basal disc, lamellae indistinct over base. Aperture subcircular with parietal concavity, broad, well-developed abapical spout. Columella weakly convex. Outer lip not preserved. Columellar callus forming broad rim, detached from base, parietal callus absent. Peristome duplex. No umbilicus.

Discussion. This species is superficially reminiscent of Anguloacrilla kimakowiczi ( Boettger, 1896) but is not distinctly coronate and differs in its much greater number of lamellae. The close-set lamellae also distinguish Anguloacrilla lima sp. nov. from other Paratethyan congeners.

‘Acrilla’ leptoglyptamoena Sacco, 1891, from the Langhian or Burdigalian of the Colli Torinesi ( Italy), and ‘ Acrilla’ praecurrens Sacco, 1891, from the Rupelian of Carcare ( Italy), are reminiscent of Anguloacrilla lima sp. nov. in their densely spaced lamellae but lack any shoulder and their spiral sculpture equals the axial lamellae in strength (see Sacco 1891: pl. 2, figs 60–61, Ferrero Mortara et al. 1984: pl. 6, fig. 1). This species resembles Amaea species, but has a narrow, but distinct subsutural ramp and shoulder, absent in Amaea .

Paleoenvironment. Unknown.

Distribution in Central Paratethys. Badenian (Middle Miocene): Făget Basin: Lăpugiu de Sus ( Romania).

Genus Cirsotrema Mörch, 1852

Type species. Scalaria varicosa Lamarck, 1822 View in CoL , by monotypy. Present-day, Indo-West Pacific.

Diagnosis. “ Thick, massive shell. Size often quite large; turriculate, conical shape; slowly increasing, more or less gradate spire; acute, multispiral protoconch; convex, superimposed or subjoined whorls, with very deep, but generally invisible, suture; ornamentation composed of powerful axial lamellae, laminated or even frondose, more or less reflexed, depending on the species, sometimes almost joined and interspersed with large, even more prominent varicose veins; they are often spiny at the back; near the anterior suture, these lamellae form a retrocurrent fold which borders the suture almost continuously, and to which is welded - in the interval between two consecutive lamellae - an anterior deviation of the lamella of the preceding whorl, so that the whole completely masks the sutures; the whorl and lamellae are, moreover, adorned with spiral cords in the interspaces. Last whorl equal to a quarter - or at most a third - of the total height, slightly convex at the base, whose periphery is marked by an almost continuous cord, formed by a retrocurrent “ spike ” or “ buttress ” which is detached from the posterior part of each lamella; sometimes, this buttress is reduced to a simple thickening of the lamella, but it is never completely fused to the preceding lamella; the basal disc, between the discontinuous cordon and the imperforate center, is radiated by the sinuous lamina of the lamellae, which flatten considerably as they widen. Aperture approximately circular, with a very thick , continuous peristome, generally with a posterior auricle, and located in a vertical plane. ” (translated and slightly modified from Cossmann 1912: 50).