Amaea hoernesi ( De Gregorio, 1889 )

Amaea hoernesi ( De Gregorio, 1889)

Figs 2K View FIGURE 2 , 11A–B View FIGURE 11

Scalaria amoena Phil.—Hörnes 1856: 479 , pl. 46, figs 11a–b [non Amaea amoena ( Philippi, 1844) ].

* [ Scalaria cancellata ] Var. Hörnesi— De Gregorio 1889: 9 [nov. nom. pro Scalaria amoena in Hörnes 1856: pl. 46, figs 11ab].

[ Scalaria View in CoL ] A [crilla]. amoena var. mioBronni Sacc.— Sacco 1891: 64.

Scalaria (Acrilla) amoena Phil. var. subcancellata d’Orb.— Boettger 1902: 86 .

Scala (Acrilla) amoena hoernesi Pant.— Sieber 1956: 313 [non Scalaria (Cirsotrema) hornesi

Pantanelli, 1886].

A [maea]. (A [crilla].) amoena subreticulata (Orb.) — Sieber 1958: 129 [non Amaea subreticulata (d’Orbigny, 1852) ].

Scala (Acrilla) amoena subreticulata d’Orb. — Csepreghy-Meznerics 1969a: 22, pl. 4, fig. 3 [non Amaea subreticulata (d’Orbigny, 1852) View in CoL ].

Scala (Acrilla) amoena subreticulata d’Orb. — Csepreghy-Meznerics 1969b: 74, pl. 1, fig. 30 [non Amaea subreticulata (d’Orbigny, 1852) View in CoL ].

Acrilla (Acrilla) subreticulata (d’Orbigny) — Bałuk 1975: 175, pl. 21, figs 5–7 [non Amaea subreticulata (d’Orbigny, 1852) View in CoL ]. Acrilla subreticulata (d‘Orbigny, 1852) — Bałuk 2006: 203, pl. 10, fig. 6 [non Amaea subreticulata (d’Orbigny, 1852) View in CoL ].

non Acrilla View in CoL mio-Bronni Sacco— Cossmann 1912: 57, pl. 3, figs 10–11 [= Amaea sp. ].

Type material. Holotype, NHMW 1858 View Materials /0015/0196, SL: 16.5 mm, MD: 6.0 mm, Baden ( Austria), illustrated in Hörnes (1856: pl. 46, figs 11a–b), Figs 2K View FIGURE 2 , 11A View FIGURE 11 1 –A View FIGURE 1 2 View FIGURE 2 .

Illustrated material. NHMW 1867/0019/0256, SL: 13.7 mm, MD: 6.3 mm, CoŞteiu de Sus ( Romania), Figs 11B View FIGURE 11 1 –B View FIGURE 1 2 View FIGURE 2 .

Revised description. Shell medium-sized, moderately slender (apical angle ~22°). Protoconch unknown. Teleoconch of more than nine relatively low whorls. Whorl profile strongly convex. Early teleoconch whorls with straight-sided to weakly concave subsutural ramp, weakly angled periphery placed mid-whorl; later whorls evenly convex. Suture moderately impressed, linear. Sculpture of delicate, slightly raised orthocline to weakly prosocline lamellae, not aligned axially (~20: ~40: ~45). Lamellae slightly alate and bent in adapertural direction. Spiral sculpture on early whorls of two weak spiral cords on subsutural ramp and three primary spirals along periphery. Secondary spiral cords intercalated on later whorls. Spiral sculpture of later whorls only slightly less raised than lamellae, forming fine, regularly cancellate sculpture. Faint spiral threads and delicate growth lines in interspaces. Last whorl attaining about 37% of total height, with about seven to nine spiral cords, secondary cords typically weaker but may attain equal strength to primaries, basal angulation delimited by moderately prominent, narrow peribasal cord, delicate lamellae and spiral threads on basal disc. Aperture ovate with parietal concavity. Outer lip and peristome not preserved. Columellar callus and parietal callus forming broad, relatively thick rim, slightly detached from base. Umbilicus reduced to very narrow chink.

Discussion. Scalaria (Acrilla) miobronni Sacco, 1891 , from the Tortonian of S. Agata ( Italy), corresponds in all features with the Paratethyan shells. Therefore, we treat it as subjective junior synonym of Amaea hoernesi ( De Gregorio, 1889) . A specimen from the Serravallian of Saubrigues ( France, illustrated by Cossmann (1912: pl. 3, figs 10–11) has a weak shoulder angulation and slightly wider cancellate sculpture and is excluded herein from synonymy. Amaea subreticulata (d’Orbigny, 1852) from the Early Miocene of the North Sea, as illustrated by Glibert (1952: pl. 7, fig. 1), has much weaker lamellae that are not elevated over the spiral cords. Amaea hoernesi was confused by Hörnes (1856) with Amaea amoena ( Philippi, 1844) , from the Oligocene of the North Sea, which differs in its less incised suture and less convex whorls. The Oligocene species has broad spiral cords, which are separated by narrow interspaces ( Janssen 1978: pl. 13, fig. 65). Amaea bronni ( Seguenza, 1876) , from the Mediterranean Pliocene, differs in its higher lamellae and spiral cords and the weakly coronate shoulder (see Landau et al. 2006: pl. 14, figs 2, 3; Chirli 2009: pl. 17, figs 12–17, pl. 18, fig. 1). For a separation from Amaea grunerti sp. nov. see below.

Paleoenvironment. Unknown.

Distribution in Central Paratethys. Badenian (Middle Miocene): Korytnica Basin: Korytnica ( Poland) ( Bałuk 2006); Vienna Basin: Baden ( Austria) (Hörnes 1856); Făget Basin: CoŞteiu de Sus ( Romania) ( Boettger 1902).

Proto-Mediterranean Sea. Tortonian (Late Miocene): Po Basin: S. Agata ( Italy) ( Sacco 1891).