Prodalmanniini Aczél, 1956

Tribe Prodalmanniini Aczél, 1956

Aczél, 1956: 3.

Type genus: Prodalmannia Bezzi, 1929 .

D i a g n o s i s. Ocelli present or absent; frontal setae lacking; antennae very short, pedicel with deep incision (P); face with short, partly fused antennal grooves; epistome very high, weakly sclerotized (P); wing with subcostal vein entire and reaching costal vein at acute angle (?A); costal vein with humeral and subcostal break (P); abdomen with sternites 1 and 2 fused ( SA with other Pyrgotinae ) with visible seam (P); male genitalia: phallic guide not developed (P); epandrium densely setose dorsally (P); inner surstylus with prensisetae (2–4 thickened setae) (P); female: oviscape on ventral side with a tongue-like comb of thickened setae (A); eversible membrane with one pair of taeniae (P); aculeus moderately large, twice as long as oviscape width at aperture (P), with setulose sternite 6 (P) and well developed anal slit (P); ventral receptacle scepter-like ( SA with other Pyrgotinae ).

This tribe is believed to be represented by a single species of its type genus, Prodalmannia variabilis Bezzi, 1929 , which occurs only in Australia. Another undescribed new species of an undescribed new genus is known to me from Mt. Magnet , Western Australia, in the ANIC .

T a x o n o m i c p o s i t i o n. Prodalmanniini have fused sternites 1+2 ( synapomorphy with the Toxurini and Pyrgotini) and thus belong in the subfamily Pyrgotinae , differing from the two latter tribes by having prensisetae on inner surstylus of males (setulose or microtrichose, without setae or prensisetae in the Toxurini and Pyrgotini) and aculeus with well-developed, setulose sternite 8 of females (“ventral lobes of the aculeus”) (almost entirely membranous, without setulae in the Toxopyrgotini trib. n., Toxurini and Pyrgotini) and well-developed anal slit on the dorsal side of cercal unit of female (no visible anal aperture in the Toxopyrgotini trib. n., Toxurini and Pyrgotini; rectum apparently opening within tergosternite 8 forming a cloaca).

Prodalmanniini form the basalmost branch of the Pyrgotinae , opposite to the Toxopyrgotini trib. n. + Toxurini + Pyrgotini lineage.

Genus Prodalmannia Bezzi, 1929 View in CoL

Type species: Prodalmannia variabilis Bezzi, 1929 (by original designation).

D i a g n o s i s. The genus can be distinguished from other pyrgotids by the combination of the tribal characters (see above) and the ocelli lacking ( apomorphy, apparently lost independently from most other tribes of the Pyrgotinae ); body densely and long setulose, subshining, not microtrichose ( fig. 1–3 View Fig ); high, poorly sclerotized epistome, gena as high as eye, the subocular sclerite not developed, the proboscis elongate, geniculate; the femora not swollen, without femoral organ, long and densely setulose, without short spinulae; and wing with vein R 4+5 bearing 4–5 setae basally on dorsal side and lacking pattern; aculeus wide, with narrow and pointed cercal unit.

R e m a r k s. A monotypic genus known only from Australia. (Korneyev, in preparation).