Aetheapnomyia hoffeinsorum ( Szadziewski, 1998 )
publication ID |
https://doi.org/10.11646/zootaxa.5706.2.2 |
publication LSID |
lsid:zoobank.org:pub:9F2CA2DA-4B2D-4187-AB12-898047C7FDFD |
persistent identifier |
https://treatment.plazi.org/id/03DB87C5-8D76-8C06-ECAA-F8B5FC95FB60 |
treatment provided by |
Plazi |
scientific name |
Aetheapnomyia hoffeinsorum ( Szadziewski, 1998 ) |
status |
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Aetheapnomyia hoffeinsorum ( Szadziewski, 1998) View in CoL
( Figs 1–4 View FIGURE 1 View FIGURE 2 View FIGURE 3 View FIGURE 4 )
Aedes ( Finlaya) hoffeinsorum Szadziewski, 1998: 235 View in CoL ( ♂, ♀; Baltic amber). Aetheapnomyia hoffeinsorum View in CoL of Harbach & Greenwalt, 2012: 33 (comb. nov., gen. nov.); Giłka et al. 2021: 259 (checklist of fossil mosquitoes); Harbach 2024, 2025 (taxonomy).
Material examined. Adult male, complete specimen (visible only from lateral side, visibility of head and genitalia limited by resin layers between which the specimen is located; right midleg attached, remaining legs present but detached, lying next to specimen), preserved in a piece of amber 46 × 20 × 17 mm (Eocene, Priabonian, 33.9–37.8 Mya), Zarechnoye District, Rovno Region, Ukraine ( SIZK Z-9; Fig. 1A–C View FIGURE 1 ). Syninclusions: Coleoptera incertae sedis, 1 specimen ; Hymenoptera incertae sedis, 1 specimen.
Description. Adult male. Colour: Body, mouthparts and legs brown to dark brown or black; wing membrane uniformly coloured, without spots; scales of thorax and wing pale, scales darker on mouthparts, legs and abdomen.
Size: Body length, excluding proboscis, 4.4 mm, body + proboscis 5.9 mm, thorax + abdomen 3.9 mm.
Antenna ( Figs 1C View FIGURE 1 , 2A–C View FIGURE 2 ):Length including pedicel 2.14 mm; flagellum 1.99 mm, comprised of 13 flagellomeres, flagellomeres 12 and 13 longest (0.39 and 0.58 mm, respectively), their combined length slightly less than total length of preceding flagellomeres (ratio 0.95); flagellar whorls well-developed, flagellomeres 1–12 each with whorl of many long setae, flagellomere 13 with whorl of several shorter setae and conical apical prolongation; antenna about 1.43 × length of proboscis.
Proboscis ( Figs 1C View FIGURE 1 , 2A, D View FIGURE 2 ): Length 1.50 mm, longer than maxillary palpus (ratio 1.30), flattened dorsoventrally and gradually broadened toward apex; labella of typical aedine form, with well-defined proximal and distal sclerites, proximal sclerite crescentic in lateral view, dorsal sclerite slightly longer than proximal sclerite, shaped like convex surface of bowl of a spoon, both sclerites largely covered with setae.
Maxillary palpus ( Fig. 2A, E View FIGURE 2 ): Length 1.15 mm, shorter than proboscis (ratio 0.77), with 5(?) palpomeres (possibly only 4, see Discussion) combined length of palpomeres 1–3 882 μm, about 0.59 length of proboscis; palpomere 4 (209 μm) more than 3 times longer than palpomere 5 (67 μm), ratio 3.12; palpomere 1 (= palpifer?, see Discussion) short, somewhat bulbous; palpomeres 2–4 slender, each gently broadening toward apex; palpomere 5 greatly reduced, ovoid with rounded apex, bearing up to 5 long setae distally.
Thorax ( Figs 1C View FIGURE 1 , 2A View FIGURE 2 ): Integument dark brown (some areas obscured by milky amber); paratergite with pale scales; postspiracular, prealar, upper and lower mesokatepisternal and upper and lower mesepimeral setae present; some pale scales on upper mesokatepisternal, lower mesokatepisternal, and upper and lower mesepimeral areas.
Wing ( Fig. 3A, B View FIGURE 3 ): Length 2.60 mm (base to tip), 2.29 mm (arculus to tip), maximum width 0.63 mm; scales spatulate, slender, apically rounded or slightly apically triangulate, apices minutely pectinate; cells R 2 and R 3 distinctly shorter than vein R 2+3 (R 2 /R 2+3 = 0.80, R 3 /R 2+3 = 0.90); arrangement of veins in area of radiomedial and mediocubital crossveins as shown in Fig. 3 View FIGURE 3 ; mediocubital crossvein [base of M 3+4 of Harbach & Knight 1980] unusually short; cubitus posterior (plical vein of Belkin 1962) developed as strongly as cubitus anterior, with scales at least on proximal part; anal vein (1A) ending 1/3 (0.35) of distance between intersection of mediocubital crossvein and cubitus and base of M 3+4 toward apex of wing.
Legs ( Figs 1C View FIGURE 1 , 4A–D View FIGURE 4 ): Entirely dark-scaled; fore- and midungues borne well before apex of tarsomere 5, long, curved, unequal on foreleg (lengths 90 μm and 120 μm) and midleg (longer unguis 110 μm, shorter unguis unmeasurable), short and equal on hindleg (~60 μm); all ungues simple, without teeth. Lengths of segments of legs in Table 2.
Abdomen ( Fig. 1C View FIGURE 1 ): Laterotergite of segment I apparently without scales, with setae only; tergum II with a few basolateral pale scales; terga III‒VIII apparently without pale bands, entirely dark-scaled; scaling of sterna uncertain, but some dark scales evident on lateral areas of sterna III‒VI.
Genitalia ( Fig. 4E, F View FIGURE 4 ): Observable in dorsolateral view (in part obscured by layer of milky amber); gonocoxite elongate, length ~300 µm; gonostylus slender, slightly arched, about half length of gonocoxite, with long, slender terminal gonostylar claw; claspette moderately long with narrow columnar stem and terminal flattened filament (see Szadziewski 1998: fig. 3).
Scales ( Fig. 4G–Q View FIGURE 4 ): Variable in shape, depending on location on wing veins ( Fig. 4G–J View FIGURE 4 ), proboscis ( Fig. 4K View FIGURE 4 ), maxillary palpus ( Fig. 4L View FIGURE 4 ), thorax ( Fig. 4M View FIGURE 4 ), legs ( Fig. 4N, O View FIGURE 4 ) or abdomen ( Fig. 4P, Q View FIGURE 4 ); ranging from broadly spatulate (twice as long as wide at most) with apex slightly triangulate or rounded, to slender (several times as long as wide) with apex more or less convex; apices pectinate.
SIZK |
Schmaulhausen Institute of Zoology |
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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Aetheapnomyia hoffeinsorum ( Szadziewski, 1998 )
Zakrzewska, Marta, Perkovsky, Evgeny E., Harbach, Ralph E. & Giłka, Wojciech 2025 |
Aedes ( Finlaya ) hoffeinsorum
Gilka, W. & Harbach, R. E. & Perkovsky, E. E. 2021: 259 |
Harbach, R. & Greenwalt, D. 2012: 33 |
Szadziewski, R. 1998: 235 |