Biblis aganisa Boisduval, 1836

Biblis aganisa Boisduval, 1836 View in CoL is a species distinct from Biblis hyperia (Cramer, 1779)

Genetic diversification between the nominotypical Biblis (Biblis) hyperia (Cramer, 1779) ( type locality St. Thomas) and Biblis (Biblis) hyperia aganisa Boisduval, 1836 ( type locality inferred as Mexico) is at the level of that between two species from the subgenus Vila : Biblis (Vila) azeca (Doubleday, [1848]) and Biblis (Vila) eueidiformis (Joicey & Talbot, 1918) (Fig. 38). The COI barcodes of the two taxa differ by 4.6% (30 bp). The nominotypical B. hyperia is characterized by more prominent dorsal forewing marginal bands than B. b. aganisa . Taken together, these arguments suggest that Biblis aganisa Boisduval, 1836 reinstated status is a species distinct from Biblis hyperia (Cramer, 1779) .

The following taxa are junior subjective synonyms: Catacore Dillon, 1948 of Diaethria Billberg, 1820 ; Mesotaenia Kirby, 1871 with Orophila Staudinger, 1886 of Perisama Doubleday, 1849 ; and Paulogramma Dillon, 1948 of Catagramma Boisduval, 1836

Genomic sequencing and analysis of the type species of available genus-group names of the subtribe Callicorina Orfila, 1952 resulted in a fully resolved confident phylogeny (Fig. 39) that revealed close relationships between some of them. For instance, even the most distinctive of the type species for available genus-group names in the Diaethria Billberg, 1820 ( type species Papilio clymena Cramer, 1775 ) clade sometimes separated in the monotypic genus Catacore Dillon, 1948 ( type and the only species Catagramma kolyma Hewitson, 1851 ) clusters closely with Diaethria . The COI barcodes of Diaethria and Catacore type species are only 6.7% (44 bp) different. In agreements with Wahlberg (2019a), we treat Catacore Dillon, 1948 as a junior subjective synonym of Diaethria Billberg, 1820 . Next, the three genera Perisama Doubleday, 1849 ( type species Catagramma bomplandii Guérin-Méneville, [1844] ), Mesotaenia Kirby, 1871 ( type species Callitaenia doris C. & R. Felder, 1861, a subspecies of Catagramma vaninka Hewitson, 1855 )

and Orophila Staudinger, 1886 ( type species Cybelis campaspe Hewitson, 1869 , a subspecies of Cybdelis cardases Hewitson, 1869 ) cluster closely together (Fig.

39). Their COI barcodes also indicate a close relationship, e.g., the type species of Mesotaenia and

Perisama are only 6.2% (41 bp) different. Therefore, Fig. 39. Callicorina genera: Lucinia (black), Haematera (red: subgenus Antigonis , and cyan),

we agree with Wahlberg (2019a) and confirm that Catagramma (olive & pink), and Callicore (purple & Mesotaenia Kirby, 1871 and Orophila Staudinger, 1886 yellow: subgenus Diaethria , magenta & green:

are junior subjective synonyms of Perisama Doubleday , subgenus Perisama , and blue).

1849. Furthermore, the genomic tree confirms the close relationship between Paulogramma Dillon, 1948 ( type species Nymphalis pyracmon Godart, [1824] ) and Catagramma Boisduval, 1836 ( type species Catagramma hydaspes Boisduval, 1836 , which is a junior subjective synonym of Nymphalis pygas Godart, [1824] ). The reasons why Freitas et al. (2014) resurrected Paulogramma in favor of the older name Catagramma that they did not use are unclear. Their tree ( Freitas et al. 2014: Fig. 3) shows " Callicore pygas ", the current name for the type species of Catagramma , inside Paulogramma as they define it. Furthermore, Wahlberg (2019a) lists Paulogramma in synonymy with Catagramma . Therefore, we confirm that Paulogramma Dillon, 1948 is a junior subjective synonym of Catagramma Boisduval, 1836 . Finally, we confirm that Lucinia Hübner, [1823] belongs to Callicorina (Fig. 39).