Homoneura ( Homoneura ) beckeri (Kertesz, 1900)

Key to all species of the Homoneura ( Homoneura) beckeri View in CoL group in Oriental Region

(Figures for previously described species can be found in the cited references)

1. At least abdominal tergites 1–2 entirely black............................................................... 2

- Abdominal tergites 1–2 mostly yellow..................................................................... 8

2. At least one of abdominal tergites 1–5 with yellow markings................................................... 3

- Abdominal tergites 1–5 entirely deep black................................................................. 5

3. Abdominal tergites 1–4 entirely black ( Fig. 38); aedeagus with 3 pair of long sword-like apical processes ( Figs. 70–71) H. ( H.) ensata sp. nov.

- At least one of abdominal tergites 1–4 with yellow markings; aedeagus not as above................................. 4

4. Upper half of occiput with a large brown spot on each side, extending to top margin of eye; abdominal tergite 3 with narrow yellow anterior margin, tergites 4–5 black on posterior 1/2 and yellow on anterior 1/2; aedeagus with a pair of falciform dorsal sclerites in ventral view ( Malloch 1929)............................................... H. ( H.) occipitalis Malloch View in CoL

- Upper half of occiput usually without large brown spot on each side; abdominal tergite 3 yellow anterolaterally; tergites 4–6 each yellow with a blackish brown median spot, which is confluent with blackish brown posterior band in tergites 4–5; aedeagus not as above( Figs. 94–95)........................................................ H. ( H.) trilineata View in CoL sp. nov.

5. Abdominal tergite 6 entirely black................................................ H. ( H.) bomiensis Gao & Yang View in CoL

- Abdominal tergite 6 yellow with brown median spot.......................................................... 6

6. Face yellow with a pair of black spots; surstylus not curved in lateral view; hypandrium without ventral processes ( Sasakawa 1992)...................................................................... H. ( H.) immaculata (de Meijere) View in CoL

- Face yellow with a large brown median spot; surstylus curved in lateral view; hypandrium with a pair of broad short ventral processes............................................................................................ 7

7. First flagellomere yellow in basal half and blackish brown in apical half ( Fig. 5); mesoscutum with a pair of brownish yellow lateral stripes ( Fig. 16)......................................................... H. ( H.) dulongjiangica View in CoL sp. nov

- First flagellomere entirely blackish brown ( Fig. 4); mesoscutum without lateral stripes ( Fig. 15)...... H. ( H.) cornis View in CoL sp. nov.

8. Face with brown stripe, band or spot......................................................................9

- Face without stripe, band or spot........................................................................ 14

9. Face with one transverse band or two small horizontal spots on ventral 1/3 or near ventral margin..................... 10

- Face with a pair of large brown elliptical spots (sometimes two spots slightly confluent), occupying majority of face...... 11

10. Antennal 1st flagellomere yellow with dark apex; hypandrium slightly concave at middle; aedeagus with a pair of acuate lateral processes in ventral view ( Sasakawa 1992)........................................... H. ( H.) beckeri (Kertész) View in CoL

- Antennal 1st flagellomere entirely black; hypandrium projecting at middle; aedeagus without acuate lateral processes in ventral view ( Yang, Hu & Zhu 2002).......................................... H. ( H.) nigrantennata Yang, Hu & Zhu

11. Abdominal tergite 6 with a pair of blackish brown lateral bands ( Fig. 34)............... .. H. ( H.) baihualingensis View in CoL sp. nov.

- Abdominal tergite 6 without lateral bands................................................................. 12

12. Abdominal tergite 5 with a triangular median spot but lacking a blackish brown transverse band; protandrium with a ventral process; aedeagus consisting of a basal anterior sclerite with a pair of triangular lateral processes, a pair of basally furcated dorsal sclerites convergent at middle and a pair of hammer-shaped dorsal processes in ventral view (Shi & Yang 2009)..................................................................................... H. ( H.) convergens Shi & Yang View in CoL

- Abdominal tergite 5 with a triangular median spot confluent with a blackish brown transverse posterior band; protandrium without ventral process; aedeagus not as above............................................................. 13

13. Mesoscutum with 5 brownish yellow longitudinal stripes ( Fig. 14); surstylus consisting of 1 process; aedeagus distinctly incised, with a pair of dorsal sclerites and a pair of preapical lateral processes ( Figs. 52–56)......... H. ( H.) conica View in CoL sp. nov.

- Mesoscutum without longitudinal stripes; surstylus consisting of 2 processes; aedeagus not as above (Shi & Yang 2009)................................................................................. H. ( H.) convergens Shi & Yang View in CoL

14. Fore tibia with an incomplete brown ring at basal 1/3.................................. H. ( H.) gemmiformis View in CoL sp. nov.

- Fore tibia without brown ring........................................................................... 15

15. Antennal 1st flagellomere unicolorous.................................................................... 16

- Antennal 1st flagellomere bicolored...................................................................... 19

16. Antennal 1st flagellomere brownish yellow; aedeagus with a pair of acuate lateral processes, a pair of incurved horn–like apical processes in ventral view (Shi & Yang 2009).................................... H. ( H.) yinggelingica Shi & Yang

- Antennal 1st flagellomere entirely black; aedeagus not as above................................................ 17

17. Wing immaculate ( Fig. 31); protandrium with a long narrow furcated ventral process; surstylus broad hairy digitiform; aedeagus without distinct apical incision, consisting of a pair of dorsal sclerites, with a pair of lateral median teeth, a pair of ventral teeth and a pair of dorsal teeth ( Figs. 81–85)........................................ .. H. ( H.) longiprocessa sp. nov.

- Wing with a narrow brown stripe or spot on dm-cu; male genitalia not as above................................... 18

18. Mesoscutum with brown stripe; tergite 6 with a small brown median spot; surstylus with a knife-shaped process in lateral view; aedeagus with a pair of acuate triangular ventral processes and a pair of subuliform dorsal processes in ventral view (Shi & Yang 2009)..................................................................... H. ( H.) flavida Shi & Yang View in CoL

- Mesoscutum without stripe; tergite 6 without spot; surstylus consisting of a broad hairy digitiform process and a narrow curved bar-like process in lateral view; aedeagus with a pair of acuate lateral processes and a pair of wide furcated dorsoapical processes in ventral view (Shi & Yang 2009)........................................... H. ( H.) tortifurcata Shi & Yang

19. Wing with a narrow brown stripe or spot on dm-cu.......................................................... 20

- Wing immaculate, without stripe or spot.................................................................. 25

20. Abdominal tergite 6 without spot; hypandrium with a pair of short bar-like ventral processes, aedeagus with a pair of dorsoapical teeth (Shi & Yang 2009)...................................................... H. ( H.) apicomata Shi & Yang View in CoL

- Abdominal tergite 6 with a brown median spot; hypandrium and aedeagus not as above............................. 21

21. Tarsomere 5 entirely yellow to yellowish brown; surstylus undulating (Shi & Yang 2009.... H. ( H.) semicircularis Shi & Yang

- Tarsomere 5 partly or entirely brown to black; surstylus not undulating.......................................... 22

22. Tarsomere 5 brownish black; surstylus only with a stick-like process, slightly curved and narrow apically; aedeagus with a pair of curved subuliform processes, a pair of acuate lateral processes and a pair of dorsal teeth on dorsal sclerites (Shi & Yang 2009)........................................................................ H. ( H.) nigritarsis Shi & Yang View in CoL

- Tarsomere 5 dark yellow in basal half and brown in apical half; surstylus consisting of a small subuliform process and a digitifom process; aedeagus with 1-3 pairs of small acuate processes or teeth on dorsal sclerites.......................... 23

23. r-m before middle of discal cell ( Fig. 32); abdominal tergite 2 without spot or band ( Fig. 41); aedeagus with a pair of slender dorsal sclerites excurved at apex, and a pair of long coniform lateral processes at base ( Figs. 89–90).. H. ( H.) pilifera View in CoL sp. nov.

- r-m at middle of discal cell; abdominal tergite 2 with spot or band; aedeagus not as above........................... 24

24. Protandrium with a long T-shaped ventral process; aedeagus with dorsal sclerites curved in lateral view (Shi & Yang 2009)................................................................................. H. ( H.) latissima Shi & Yang View in CoL

- Protandrium with a short V-shaped ventral process; aedeagus with dorsal sclerites straight in lateral view ( Li, Li & Yang 2008)......................................................................... H. ( H.) yunnanensis Li, Li & Yang View in CoL

25. Surstylus consisting of 3 processes in lateral view; hypandrium with a pair of long apodemes and a small apodeme at median incision; aedeagus with 3 pairs of preapical dorsal teeth ( Figs. 76–80)......................... H. ( H.) latizona View in CoL sp. nov.

- Surstylus consisting of 1 process in lateral view; hypandrium and aedeagus not as above............................ 26

26. Abdominal tergites 3–5 without median spot ( Fig. 35); hypandrium without hypandrial apodeme; gonopod incurved at apex in ventral view; aedeagal apodeme short ( Figs. 50–51)..................................... .. H. ( H.) brevituba View in CoL sp. nov.

- Abdominal tergites 3–5 each with a median spot; male genitalia not as above.....................................27

27. Abdominal tergites 3–5 with blackish brown posterior band confluent with triangular median spot; surstylus short claw-like, not projecting beyond tip of epandrium in lateral view; hypandrium without lobate ventral sclerites in ventral view; aedeagus with bifurcated sclerite ventrally and two pairs of spine-like processes on dorsal and ventral membranous parts ( Sasakawa 1992)......................................................................... H. ( H.) fasciventris Malloch View in CoL

- Abdominal tergites 3–5 each with a black median spot and a pair of black lateral bands; surstylus long knife-like, projecting beyond tip of epandrium in lateral view; hypandrium with a pair of lobate ventral sclerites in ventral view; aedeagus obliquely truncate at tip with a small acuate process (Shi & Yang 2009).............................. H. ( H.) zhangae Shi & Yang