Nyctiophylax ( Paranyctiophylax ) kupirah, Oláh & Johanson, 2010

Nyctiophylax ( Paranyctiophylax) kupirah , new species

Figs 23–26 View FIGURES 23–26

This species belongs in the Nyctiophylax ( Paranyctiophylax) flavus species group and is most similar to N. padangensis from North Sumatra, but differs by its triangular, not spatulate cerci; ventral paraproctal processes each with shorter ventrad-curving apical part; gonopods each with well-developed ventrobasal “elbow;” the shorter and basally curved ventroapical median lobe of sternite IX; and the presence of a pair of strong, anterad-directed spines and an ovoid phallotremal sclerite in the phallic apparatus.

Male. Body uniformly pale brown. Maxillary palp formula IV-II-I-III-V, segment III originating from mesal face basal of apex of segment II. Forewings 4.7 mm long; brown, with hypertrophied longitudinal vein R and pronounced hyaline patch pattern: small, rounded spot below discoidal cell around crossvein r-m in subradial cell; and large, triangular spot along crossvein m-cu and point of basal bifurcation of M; discoidal cell and median cell present; apical forks 2, 3, 4 and 5 present; A1, A2, and A3 looping. Hind wing discoidal cell closed, median cell open; apical forks 2 and 5 present.

Spur formula 3,4,4.

Male genitalia. Sternite IX robust, triangular in lateral view with well-developed apicoventral mesal lobe having regular rectangular apex in ventral view and sinuous basal half in lateral view, shorter than ventrobasal “elbows” of gonopods. Tergite IX small, sclerotized; elongated obliquely vertical, discernible dorsal of sternite and anterior of paraproct. Segment X membranous, nearly invisible; with triangular mesal lobe in dorsal view. Cerci setose, triangular in lateral view; clavate in dorsal view; shorter than gonopods and ventral paraproctal processes. Paraproctal complex well-developed, highly sclerotized; consisting of pair of large ventral paraproctal processes curving ventrad, each bearing 4-6 setae in row on outer surface of ventradcurving apical half. Dorsal paraproctal processes forming pair of hyaline digitate processes. Gonopods elongate, slender, uniformly curving in ventral view; each with basal half broadening, forming ventrobasal “elbow;” distal half slender; apex slightly excised, located laterally. Phallic apparatus located dorsally in genitalia, fixed, guided by paraproct and segment X; middle section of phallotheca forming short tube. Phallobase modified into long, narrow, arching dorsal apodeme and short ventral apodeme in lateral view; dorsal apodeme distinctly clear-cut, arching and narrowing into spine-like process in lateral view; weakly narrowing in dorsal view. Phallicata membranous, with pair of anterad-directed, stout spines and ovoid, horizontally elongate phallotremal sclerites embedded in phallic membrane.

Holotype male: INDONESIA ( Sumatra): Bukit Barisan Selatan National Park, Barisan Mts., Way Pitias , 800 m, 17–19.vi.2008, light trap [ J. Oláh, Jr.]— 1 male (OPC, in alcohol).

Paratypes: same data as holotype— 10 males (OPC, in alcohol) .

Distribution: Indonesia ( Sumatra).

Etymology: Kupirah, from “kupirah,” elbow in Sanskrit, referring to the large, ventrobasal “elbows” of the gonopods.