Moro bloetei ( Dlabola, 1982 ), 2025

Gnezdilov, Vladimir M. & Bieman, F. M., 2025, The smallest species of the family Issidae-a new genus for Hysterodus bloetei Dlabola, 1982 (Hemiptera: Fulgoroidea), Zootaxa 5683 (3), pp. 443-450 : 445-448

publication ID

https://doi.org/10.11646/zootaxa.5683.3.7

publication LSID

lsid:zoobank.org:pub:C8F68D91-90C1-4746-855E-A377E4EE8F0F

persistent identifier

https://treatment.plazi.org/id/03E587FA-3311-FFA0-68D8-2E7C2A2FFC2F

treatment provided by

Plazi

scientific name

Moro bloetei ( Dlabola, 1982 )
status

comb. nov.

Moro bloetei ( Dlabola, 1982) , comb. nov.

( Figs 1–20 View FIGURES 1–3 View FIGURES 4–7 View FIGURES 8–13 View FIGURES 14–20 )

Hysterodus bloetei View in CoL : Dlabola 1982: 164 (sp. n., figs 273–282) Hysterodus bloetei View in CoL : Gnezdilov 2003: 32 (list) Quadriva bloetei View in CoL : Gnezdilov et al. 2004: 218 (comb. n.)

Quadriva bloetei View in CoL : Gnezdilov et al. 2014: 99 (uncertain position)

Material examined. Spain: 16♂, 13♀, Ciudad Real , 6 km NNW Ruidera, along CM 3115, N39.03122 W2.91030, 920 m, 26. V GoogleMaps .2016 ( ZIN and CCB) GoogleMaps ; 21♂, 19♀, same locality, 8. V GoogleMaps .2017 ( ZIN and CCB) GoogleMaps ; 16♂, 14♀, same locality, 17. V GoogleMaps .2018 ( CCB) GoogleMaps ; 3♂, 5♀, Ciudad Real , 8 km NNW Ruidera, near CM 3115, N39.04506 W2.94186, 880 m, 9. V GoogleMaps .2017 ( CCB) GoogleMaps ; 1♂, Ciudad Real , 1 km SW Ruidera, along N430, N38.97031 W2.89434, 870 m, 18. V GoogleMaps .2018 ( CCB) GoogleMaps ; 2♂, 2♀, Albacete, 8 km SW Ossa de Montiel, Laguna de Ruidera , N38.89160 W2.76808, 820 m, 16. V GoogleMaps .2018 ( CCB) GoogleMaps ; 3♂, same locality, 19. V GoogleMaps .2018 ( CCB) GoogleMaps ; 5♂, 5♀, Valencia, 6 km NW Villargordo del Cabriel, near Embalsa de Contreras (water reservoir), N39.55509 W1.48678, 650 m, 22. V GoogleMaps .2018 ( CCB), all leg. C.F.M. den Bieman.

Emended description. Structure ( Figs 1–6 View FIGURES 1–3 View FIGURES 4–7 ). Metope slightly elongate, lateral margins convex, with strong median carina running from its upper margin throughout postclypeus and with weak sublateral carinae running from the same point with median carina on its upper margin, but not reaching metopoclypeal suture which is strongly convex ( Fig. 3 View FIGURES 1–3 ). Upper margin of metope obtusely angulately concaved. Pedicel globular. Rostrum reaching hind coxae, 3 rd segment narrowing apically, slightly shorter than 2 nd one. Coryphe slightly transverse, 1.5 times as wide as long medially, grooved concave; anterior margin obtusely angulately convex; posterior margin concave; lateral margins keel-shaped ( Figs 1 View FIGURES 1–3 , 4, 6 View FIGURES 4–7 ). Eyes large, each eye is nearly as broad as coryphe width. Pronotum slightly shorter than coryphe medially, with strongly convex keel-shaped anterior margin and notched medially posterior margin. Paradiscal fields of pronotum narrow behind eyes. Paranotal lobes of pronotum wide. Mesonotum twice longer than pronotum medially, with weak lateral carinae; concave at scutellum. Brachypterous, fore wings reaching hind margin of tergite VI (in one female —only hind margin of tergite III) ( Figs 1, 2 View FIGURES 1–3 , 4–6 View FIGURES 4–7 ). Fore wings nearly round, spread out, with its basal parts before anterior radial vein turned downwards nearly at right angle to the rest of corium; without hypocostal plate. Clavus almost as long as whole wing, opened. Forewing vein branching sequence: R 4, furcating basally; r-m 1–4; M 2, furcating apically; m-cua 3; CuA 2, furcating apically. Pcu and first anal vein fused before middle of clavus. All veins relief. Hind wings rudimentary. Legs not flattened neither foliated. Hind tibia with a single lateral spine medially (in one male on left leg a second lateral spine present distally to the middle one) and seven spines apically ( Figs 1 View FIGURES 1–3 , 4, 6 View FIGURES 4–7 ). First metatarsomere larger than second one, elongate, with two latero-apical and five intermediate spines. Second metatarsomere with only two latero-apical spines.

Coloration ( Figs 1–7 View FIGURES 1–3 View FIGURES 4–7 ). Generally orange brown. Metope and pronotum with dense dark brown to black dots. Rostrum with black 3 rd segment. Pedicel dark below. Antenna with dark brown 3 rd segment and flagellum. Fore wings with dark brown dots and spots in cells. Femora and tibiae with brown to dark brown longitudinal stripes. Apices of leg spines, third tarsomeres, and claws black. Abdominal tergites with black spots surrounding setae. In females, gonocoxa VIII dark brown near to anterior connective lamina of gonapophyse VIII.

Male genitalia ( Figs 8–13 View FIGURES 8–13 ). Pygofer vertically elongate, narrow, weakly convex medially in lateral view ( Fig. 8 View FIGURES 8–13 ). Anal tube large, enlarged medially (in dorsal view), 1.5 times as long as wide medially, with two rounded lateral processes ( Figs 8, 13 View FIGURES 8–13 ). Anal column short, 1/6 of whole anal tube. Phallobase strongly curved, horseshoe-shaped in lateral view, weakly sclerotized apically and covered by denticles, with two hardly sclerotized plates dorso-apically bearing teeth ( Fig. 9 View FIGURES 8–13 , spl). Aedeagus not visible above phallobase, with pair of long, half of phallobase length, ventral hooks pointed apically and directed basad. Ventral phallobase lobe long and narrow, weakly concave apically ( Fig. 10 View FIGURES 8–13 ). Style with concave hind margin ( Fig. 12 View FIGURES 8–13 ). Capitulum of style on long neck ( Fig. 12 View FIGURES 8–13 ), narrow in dorsal view ( Fig. 11 View FIGURES 8–13 ), with large lateral tooth and distinct apical tooth.

Female terminalia ( Figs 7 View FIGURES 4–7 , 14–20 View FIGURES 14–20 ). Hind margin of sternite VII deeply concave medially ( Fig. 14 View FIGURES 14–20 ). Anal tube elongate, 1.6 times as long as wide medially, nearly oval, with pointed apex in dorsal view ( Fig. 17 View FIGURES 14–20 ); convex in lateral view ( Fig. 18 View FIGURES 14–20 ). Anal column short. Gonoplacs convex, without carinae ( Fig. 7 View FIGURES 4–7 ). Gonocoxa VIII with lobe-shaped hind margin ( Fig. 19 View FIGURES 14–20 ). Anterior connective lamina of gonapophyse VIII wide, with two teeth in apical group (one distinct apical tooth and one smooth tooth below) and 4–5 keeled teeth in lateral group ( Fig. 19 View FIGURES 14–20 ). Endogonocoxal process with simple narrow apical part. Posterior connective laminae of gonapophyses IX wide in ventral and lateral views, with large and convex median field covered by small spines bearing two widely spaced lobes apically and with small lateral fields covered by microsetae ( Figs 15, 16 View FIGURES 14–20 ). Distal parts of the laminae curved at obtuse angle in ventral view ( Fig. 15 View FIGURES 14–20 ). Gonospiculum bridge large, elongate in lateral view ( Fig. 16 View FIGURES 14–20 ). Diverticulum ductus with slightly elongate bulb ( Fig. 20 View FIGURES 14–20 ).

Total length. Males—2.0– 2.5 mm. Females— 2.5–2.9 mm.

CM

Chongqing Museum

V

Royal British Columbia Museum - Herbarium

ZIN

Russian Academy of Sciences, Zoological Institute, Zoological Museum

CCB

Central College, Bangalore

VI

Mykotektet, National Veterinary Institute

R

Departamento de Geologia, Universidad de Chile

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Hemiptera

Family

Lophopidae

Genus

Moro

Loc

Moro bloetei ( Dlabola, 1982 )

Gnezdilov, Vladimir M. & Bieman, F. M. 2025
2025
Loc

Quadriva bloetei

Gnezdilov, V. M. & Holzinger, W. E. & Wilson, M. R. 2014: 99
2014
Loc

Hysterodus bloetei

Gnezdilov, V. M. 2003: 32
Dlabola, J. 1982: 164
1982
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