Nezzazata isabellae Arnaud-Vanneau & Sliter, 1995

Nezzazata isabellae Arnaud-Vanneau & Sliter, 1995 View in CoL

Fig. 12 View Fig

? 1971 Nezzazata cf. convexa - Fleury, 187-188, pl. 3, figs. 1-7; middle Cenomanian, Greece.

1977 Nezzazata simplex germanica - Velić, pl. XIX, figs. 4-6; “Neocomian”, Croatia [early Hauterivian according to Velić, 2007].

1977 Nezzazata simplex - Velić, pl. XXI, figs. 9-11; Barremian, Croatia.

? 1982 Nezzazata sp. C - Altiner & Decrouez, pl. VI, fig. 26; late Aptian – Cenomanian, Turkey.

T 1995 Nezzazata isabellae n. sp. Arnaud-Vanneau & Sliter, p. 552; fig. 7; pl. 2, figs. 11-24; late? Aptian-early Albian, mid Pacific Ocean.

1995 Nezzazata isabellae - Arnaud-Vanneau & Premoli Silva, p. 206; pl. 2, figs. 1-3; “middle” Aptian- Albian, Pacific/ Japan.

2006 Nezzazata isabellae - Mancinelli & Chiocchini, p. 92; pl. 4, figs. 11-21; late Aptian-early Albian, central Italy.

2006 Nezzazata simplex - Taslı et al., fig. 6F (non E = N. simplex ); middle – late Cenomanian, Turkish Taurides.

? 2010 Nezzazata isabellae - Cavin et al., fig. 9H; late Cenomanian, Morocco.

2012 Nezzazata isabellae - Chiocchini et al., pl. 66, figs. 1-14; late Aptian with range to earliest Cenomanian, central Italy.

? 2012 Nezzazata isabellae - Ghanem et al., figs. 6a(12), 6b (6, 10, 11); late Aptian – early Albian, Syria.

2013 Nezzazata isabellae - Hfaiedh et al., fig. 12W-Z, Aa; early Aptian, Tunisia.

Non 2013 Nezzazata isabellae - Ghanem & Kuss, fig. 10(14) [=? Nezzazatinella sp. ]., fig. 11(22) [indeterminate]; early – late Albian, Syria.

? 2015 Nezzazata isabellae - Citton et al., pl. 2.1;?late Aptian – early Albian?, southern Italy.

? 2016 Nezzazata simplex germanica - Hosseini et al., fig. 11d (non e = Nezzazatinella sp. ?); Hauterivian, Iranian Zagros

2019 Nezzazata isabellae - Taslı & Solak, fig. 10(8); late Albian, Turkish Taurides.

2019 Nezzazata isabellae - Özkan & Altiner, fig. 5(5); early Aptian, Turkish Arabian Plate.

? 2020 Nezzazata sp. - Solak et al., figs. 6F, I, fig. 11D; early – late Cenomanian, Turkish Taurides.

? 2020 Nezzazata isabellae - Randazzo et al., fig. 9C-D; early Albian, Sicily.

Non 2020 Nezzazata isabellae - Mohseni & Javanmard, fig. 11(i); Cenomanian, Iranian Zagros. [=simple, trochospiral form].

2021 Nezzazata isabellae - Solak et al., figs. 3.1 - 3.14; Albian, Turkish Taurides.

2021 Nezzazata isabellae - Solak, fig. 1L-M; Albian – middle Cenomanian, Turkish Taurides.

Non 2021 Nezzazatinella cf. isabellae - Consorti & Schlagintweit, fig. 2C-D; late Cenomanian, Kosova [= Nezzazatinella ex. gr. picardi , name used in error].

2023 Nezzazata simplex - Solak & Taslı, fig. 9G, H, I; Fig. 11T View Fig ; Fig. 13R, S, T View Fig ; early Aptian - Albian, Turkish Taurides.

Taxonomy/Identity: First formally described by Arnaud-Vanneau & Sliter (1995) from a Pacific Ocean Drilling Project borehole. It is smaller than the other Cenomanian and younger species with a maximum diameter of only 0.20 mm. See also the main discussion under N. simplex . N. isabellae is also distinguished by having a relatively high trochospire compared with N. simplex and N. ex gr. gyra , by having a well-rounded equatorial and axial periphery and by having the most chambers in the final whorl (12 cf. 8-10). Other similarities and differences are enumerated in Table 3 (above).

Filkorn & Scott (2011) described a form they called “ Nezzazata sp. cf. N. isabellae ” from the late Albian of Mexico. It is larger (0.6 mm), rounded and planoconvex in comparison to N. isabellae .

Reference Images: Arnaud-Vanneau & Sliter (1995), Fig. 7 View Fig , Pl. 2, Figs. 11 View Fig -24, p. 552.

Confident Stratigraphic Range: Hauterivian – middle-late Cenomanian. Commonly recorded from the Aptian and Albian; sparse records from older or younger strata.

Uncertain Stratigraphic Range: Not applicable

Originally assigned a late Aptian - early Albian age ( Arnaud-Vanneau & Sliter, 1995). Extended to the "mid" Aptian and top Albian ( Arnaud-Vanneau & Premoli Silva, 1995; Velić 2007) and the intra- early Aptian by Husinec et al. (2009). Most records are from the Aptian and Albian ( Mancinelli & Chiocchini, 2006; Spalluto & Caffau, 2010; Hfaiedh et al., 2013; Taslı & Solak, 2019; Scott et al., 2024 – see also discussion of N. simplex ). Nonetheless, the records of Velić (1977) of “ Nezzazata simplex germanica ” from Croatia extend the range to the Hauterivian as does that of Hosseini et al. (2016) from the Iranian Zagros.

Cenomanian records include that of Taslı et al. (2006) who recorded it as Nezzazata simplex from undifferentiated middle – late Cenomanian strata in the Turkish Taurides (see also Cenomanian record of Solak, 2021). Nezzazata cf. convexa described by Fleury (1971) from middle Cenomanian of Greece is similar in overall morphology, but a somewhat larger. Křížová et al. (2024) record the species from the guerangeri zone of the late Cenomanian in north-east Italy, but without illustration, whilst an illustration from similar aged sediments from Morocco ( Cavin et al., 2010) cannot be confirmed as this species. A record from the Cenomanian of the Iranian Zagros ( Mohseni & Javanmard, 2020) appears erroneous and the species does not appear to be a component of Nezzazata -rich assemblages from the Cenomanian of the Arabian Plate.

Geographic Range: Most records are from the Mediterranean alongside the type description of the species from guyots in the Pacific. Records from the Arabian Plate are sparse/unconfirmed and the species is yet to be confirmed as present in Mexico or Tibet .