Scathophaga spurca ( Meigen, 1826 )

Scathophaga spurca ( Meigen, 1826) View in CoL

Figs 37, 112–114.

Scatomyza suilla (Fabricius,); Fallén [1819: 5]. Misidentification [see: Michelsen, O’Hara, 2014: 77–79].

spurca Meigen, 1826: 250 View in CoL ( Scatophaga View in CoL ). Type-locality: not given.

glabrata Zetterstedt, 1838: 721 ( Scatomyza View in CoL ). Type-localities: “Lapponia Norvegica borealis — Dalekarlia Sueciae” ( Norway, Sweden).

scatomyzoides Zetterstedt, 1838: 727 ( Cordylura ). Type-locality: “Lapponia Umensi... Wilhelmina... (Lappon. — meridional.)” ( Sweden).

serotina Perris, 1839: 48 ( Scatophaga ). Type-locality: “environs de Mont-de-Marsan” (Landes, France).

REMARKS. This widespread and common species was recorded in Russia (in most cases as S. suilla ) from Karelia [ Polevoi, 1997: 309; Humala, Polevoi, 2009: 72], Kanin Peninsula [ Engelmark, 1999: 158, 159], Mordovia [ MacGowan et al., 2021: 17], Yaroslavl’ Oblast [ Ovchinnikov, 2004: 422], Krasnoyarsk Krai (Taimyr Peninsula) [ Ozerov, Barkalov, 2014: 564], Yakutia [ Sychevskaya, 1972: 150; Veselkin, 1985: 75; Bagachanova et al., 2016: 783] and Far East [ Ozerov, Krivosheina, 2014: 219].

Gorodkov published a map with the distribution area of this species on the territory of the former USSR and separately on its European part [ Gorodkov, 1980: map 68, as S. suilla ], but without the names of the points.

MATERIAL EXAMINED. Altai: River Chuya, 20 km NW of Kosh-Agach ( 50.085ºN 88.404ºE), 25–27.VII.2006, Nartshuk ( 1 ♂, ZISP); Amur Oblast: Bol’shoy Never ( 53.978ºN 124.150ºE) GoogleMaps , 7.IX.1974, Gorodkov ( 7 ♂♂, 5 ♀♀, ZISP); Arkhangelsk Oblast: 70 km N Nar’yan-Mar (ca. 68.204ºN 53.627ºE) GoogleMaps , 18.VIII.1978, Gorodkov ( 4 ♂♂, 1 ♀, ZISP); 73 km NW of Nar’yan-Mar (ca. 67.981ºN 51.608ºE) GoogleMaps , 3.VIII.1978, Gorodkov ( 3 ♂♂, 1 ♀, ZISP); Arkhangelsk ( 64.546ºN 40.567ºE) GoogleMaps , 10.VIII.1967, Gorodkov ( 8 ♂♂, 3 ♀♀, ZISP); Belush’e env. ( 66.89ºN 47.61ºE) GoogleMaps , 24.VIII.1978, Gorodkov ( 2 ♂♂, ZISP); Kanin Nos ( 68.656ºN 43.279ºE) GoogleMaps , 17.VII.1970, Gorodkov ( 4 ♂♂, 3 ♀♀, ZISP); Kargopol’ ( 61.503ºN 38.971ºE) GoogleMaps , 13.VII.1982, Gorodkov ( 1 ♀, ZISP); Kolguev I., Bugrino ( 68.782ºN 49.309ºE) GoogleMaps , 20–21.VIII.1970, Gorodkov ( 3 ♂♂, 3 ♀♀, ZISP); Lomonosovo ( 64.229ºN 41.732ºE) GoogleMaps , 31.VIII.1978, Gorodkov ( 1 ♂, ZISP); Mezen’ ( 65.85ºN 44.24ºE) GoogleMaps , 25 and 27.VIII.1978, Gorodkov ( 3 ♂♂, 2 ♀♀, ZISP); Nar’yan-Mar ( 67.636ºN 53.031ºE) GoogleMaps , 28.VIII.1970, 4, 7 and 21.VIII.1978, Gorodkov ( 5 ♂♂, 3 ♀♀, ZISP); Nizhnyaya Pesha ( 66.751ºN 47.760ºE) GoogleMaps , 22.VIII.1978, Gorodkov ( 1 ♂, 1 ♀, ZISP); Onega ( 63.922ºN 38.085ºE) GoogleMaps , 20.VII.1996, Gorodkov ( 2 ♂♂, ZISP); Solovetskiy I. ( 65.080ºN 35.686ºE) GoogleMaps , 27.VII.1975, Gorodkov ( 1 ♂, ZISP); Solvychegovsk env. ( 61.327ºN 46.918ºE, 61.342ºN 46.913ºE) GoogleMaps , 13–14.VIII.2010, D. Gavryushin ( 6 ♂♂, 8 ♀♀, ZMUM); same place , 2.IX.1981, Gorodkov ( 1 ♂, 2 ♀♀, ZISP); Velikovisochnoe ( 67.255ºN 52.0328ºE) GoogleMaps , 18.VIII.1978, Gorodkov ( 2 ♂♂, 3 ♀♀, ZISP); Bashkiria: Bashkirskiy Nature Reserve ( 53.420ºN 57.935ºE) GoogleMaps , 7.VIII.1975, Gorodkov ( 4 ♂♂, 2 ♀♀, ZISP); Beloretsk env. ( 54.051ºN 58.268ºE, 54.004ºN 58.463ºE, 53.973ºN 58.344ºE) GoogleMaps , 7–10.VIII.2012, D. Gavryushin ( 2 ♂♂, 1 ♀, ZMUM); Ufa env. ( 54.776ºN 56.068ºE) GoogleMaps , 18.VIII.1975, Gorodkov ( 2 ♂♂, 1 ♀, ZISP); Buryatia: Barguzinsky Nature Reserve (ca. 54.35ºN 109.51ºE) GoogleMaps , 20.VII.1965, Negrobov ( 2 ♂♂, 1 ♀, ZISP); Khasura ( 50.327ºN 103.282ºE) GoogleMaps , 1.VII.1971, V. Richter ( 1 ♂, ZISP); Tankhoy ( 51.554ºN 105.118ºE) GoogleMaps , 10–17. VI .1980 GoogleMaps , Barkalov ( 1 ♂, 1 ♀, ZISP); Chukotka: Markovo   GoogleMaps (ca. 64.680ºN 170.412ºE), 18.VIII.1966, Gorodkov ( 10 ♂♂, 8 ♀♀, ZISP); Irkutsk Oblast: Padun ( 56.293ºN 101.711ºE) GoogleMaps , 24. VI .1956 GoogleMaps , Monchadskiy ( 13 ♂♂, 8 ♀♀, ZISP); Kamchatka Krai: Kozyrevsk   GoogleMaps ( 56.048ºN 159.870ºE), 7.IX.1969, Gorodkov ( 2 ♂♂, 1 ♀♀, ZISP); Commander Islands , Mednyy I. ( 54.709ºN 167.709ºE) GoogleMaps , 29.VII.1932, I. Barabash ( 2 ♂♂, ZISP); Esso ( 55.927ºN 158.704ºE) GoogleMaps , 4 and 5.VIII.1978, A. Zinov’ev ( 2 ♂♂, ZISP); Commander Islands, Preobrazhenskoe ( 54.790ºN 167.578ºE) GoogleMaps , 5.IX.1959, Gorodkov ( 2 ♂♂, 1 ♀, ZISP); Karelia: Gridino ( 65.920ºN 34.669ºE) GoogleMaps , 11.IX.1984, Gorodkov ( 1 ♀, ZISP); Kartesh ( 66.33ºN 33.64ºE) GoogleMaps , 18 and 21.VII.1975, Gorodkov ( 4 ♂♂, 3 ♀♀, ZISP); Keret’ ( 66.190ºN 32.884ºE) GoogleMaps , 8.IX.1984, Gorodkov ( 1 ♂, 1 ♀, ZISP); Pudozh ( 66.804ºN 36.565ºE) GoogleMaps , 17 and 19.VII.1982, Gorodkov ( 5 ♂♂, 2 ♀♀, ZISP); Khabarovsk Krai: Bulgino env. ( 59.371ºN 143.147ºE) GoogleMaps , 29.VIII.1987, Gorodkov ( 2 ♂♂, 1 ♀, ZISP); Khakassia: W Sayan pass ( 51.717ºN 89.882ºE), 2100 m GoogleMaps , 9– 12.VII.2017, N. Vikhrev ( 6 ♂♂, 1 ♀, ZMUM) Komi: Blagoevo ( 63.41ºN 47.95ºE) GoogleMaps , 12.VII.1996, Gorodkov ( 1 ♂, ZISP); Ust-Tsilma ( 65.440ºN 52.153ºE) GoogleMaps , 9.VIII.1978, Gorodkov ( 2 ♂♂, ZISP); Yaksha ( 61.823ºN 56.821ºE) GoogleMaps , 12.VIII.1989, Gorodkov ( 10 ♂♂, 4 ♀♀, ZISP); Krasnoyarsk Krai: Baykit ( 61.68ºN 96.38ºE) GoogleMaps , 23– 25.VIII.1972, Gorodkov ( 16 ♂♂, 10 ♀♀, ZISP); Dudinka ( 69.404ºN 86.182ºE) GoogleMaps , 2–3.VII.1967, Gorodkov ( 2 ♂♂, ZISP); Igarka ( 67.457ºN 86.598ºE) GoogleMaps , River Yenisei , 30. VI . and 1.VII. 1967 GoogleMaps , Gorodkov ( 4 ♂♂, ZISP); Khatanga   GoogleMaps ( 71.981ºN 102.483ºE), 25.VIII.1971, Gorodkov ( 13 ♂♂, 8 ♀♀, ZISP); Turuchansk ( 65.797ºN 87.958ºE) GoogleMaps , 27–28. VI .1967 GoogleMaps , Gorodkov ( 6 ♂♂, 4 ♀♀, ZISP); Leningrad Oblast: Luga   GoogleMaps (ca. 58.73ºN 29.84ºE), 15.VII.1953, 16.VIII.1955, A. Stackelberg ( 1 ♂, 2 ♀♀, ZISP); Rakovichi ( 58.650ºN 29.836ºE) GoogleMaps , 7 and 31. V., 13–15. VI .1897 GoogleMaps , G. Pleske ( 5 ♂♂, 2 ♀♀, ZISP); Saint Petersburg ( 59.934ºN 30.359ºE), 29.VIII.1995, Gorodkov ( 6 ♂♂, 1 ♀, ZISP); Tolmachevo ( 58.856ºN 29.895ºE) GoogleMaps , 11. VI . and 7.VII.1935 GoogleMaps , Rohdendorf ( 5 ♂♂, 2 ♀♀, ZISP); Yashchera   GoogleMaps ( 58.894ºN 29.820ºE), 7. VI .1958 GoogleMaps , A. Stackelberg ( 1 ♂, ZISP); Lipetsk Oblast: Donskoe   GoogleMaps ( 52.618ºN 38.978ºE), 29.VIII.1978, Tryapitsyn ( 1 ♂, ZISP); Magadan Oblast: Chaybukha ( 61.80ºN 160.413ºE) GoogleMaps , 1.IX.1987, Gorodkov ( 2 ♂♂, 7 ♀♀, ZISP); Evensk ( 61.916ºN 159.234ºE) GoogleMaps , 9.IX.1987, Gorodkov ( 1 ♂, ZISP); Gizhiga ( 61.948ºN

127 128

129

160.377ºE), 5.IX.1987, Gorodkov ( 1 ♀, ZISP); Sokol ( 59.919ºN 150.752ºE), 25.VIII.1966, Gorodkov ( 4 ♂♂, 3 ♀♀, ZISP); Moscow and Moscow Oblast: Bittsa ( 55.641ºN 37.570ºE), 24– 31.VII.1936, B. Rohdendorf ( 5 ♂♂, 1 ♀, ZMUM); Dmitrov env. ( 56.316ºN 37.725ºE), 8–15.IX.2006, 19. V., 21. VI., 8 and 10.VII., and 2.IX.2007, 18. V.2010, N. Vikhrev ( 13 ♂♂, 4 ♀♀, ZMUM); Golitsyno ( 55.649ºN 37.011ºE), 21. V.1977, 21. V.1978, 31. V. and 4–7.IX.1981, 5. VI.1982, A. Shatalkin ( 4 ♂♂, 4 ♀♀, ZMUM); Izmaylovo ( 55.800ºN 37.832ºE), 18. VI.1983, A.L. Ozerov ( 1 ♂, ZMUM); Zvenigorod env ( 55.700ºN 36.722ºE), 24. V.1981, A.L. Ozerov ( 4 ♂♂, 1 ♀, ZMUM); Murmansk Oblast: Aleksandrovsk [= Polyarnyy] ( 69.198ºN 33.456ºE), IX.1923, Fridolin ( 1 ♀, ZISP); same place, 24–25.VIII.1923, V. Kuznetsov ( 2 ♂♂, 1 ♀, ZISP); Lake Vudyavr Basin (ca. 67.646ºN 33.644ºE), 18 and 23– 24. VI.1930, Fridolin ( 3 ♂♂, 1 ♀, ZISP); Dal’nie Zelentsy ( 69.117ºN 36.062ºE), 6 and 10.VIII.1981, 2.IX.1984, Gorodkov ( 7 ♂♂, 1 ♀, ZISP); Khibiny Station ( 67.673ºN 33.212ºE), 4.IX.1928, Cheburova ( 1 ♀, ZISP); Kirovsk env. ( 67.608ºN 33.661ºE), 6 and 9.VII.1975, Zinov’ev ( 1 ♂, 1 ♀, ZISP); Kovdor ( 67.563ºN 30.477ºE), 21.VIII.1995, Gorodkov ( 7 ♂♂, 4 ♀♀, ZISP); Krasnoshchel’e ( 67.351ºN 37.045ºE), 18.VIII.1981, Gorodkov ( 1 ♂, 2 ♀♀, ZISP); Sonostrov ( 66.172ºN 34.233ºE), 11.IX.19841, Gorodkov ( 1 ♂, ZISP); Sosnovka ( 66.5073ºN 40.583ºE), 23.VIII.1981, Gorodkov ( 10 ♂♂, 2 ♀♀, ZISP); Tumannyy ( 68.883ºN 35.693ºE), 4.VIII.1981, Gorodkov ( 3 ♂♂, 1 ♀, ZISP); Varzuga ( 66.394ºN 36.596ºE), 16.VIII.1995, Gorodkov ( 3 ♂♂, 1 ♀, ZISP); Novgorod Oblast: Novgorod env. ( 58.486ºN 31.284ºE), 1.X.1978, Gorodkov ( 1 ♂, 2 ♀♀, ZISP); Ramen’e ( 58.374ºN 33.430ºE), Okulovka env., 4. VI.1988, Gorodkov ( 4 ♂♂, 1 ♀, ZISP); Novosibirsk Oblast: Koynikha ( 54.568ºN 83.252ºE), 13.VIII.1977, Gorodkov ( 2 ♂♂, 1 ♀, ZISP); Primorsky Krai: Terney env. ( 45.043ºN 136.634ºE), 8.X.2016, M. Sergeev ( 1 ♀, ZMUM); Ryazan’ Oblast: Kasimov env. ( 54.969ºN 41.327ºE), 21–24.VII.2013, N. Vikhrev ( 1 ♂, ZMUM); Sakhalin Oblast: Kunashir I., Kurilsky Nature Reserve, cordon Andreevsky ( 43.54ºN 145.37ºE), 6–8.VII.2014, I.A. Gomyranov ( 1 ♂, ZMUM); Kunashir I., Sernovodsk ( 43.912ºN 145.641ºE), 8. VI.1968, V. Richter ( 10 ♂♂, 8 ♀♀, ZISP); Kunashir I., Yuzhno-Kuril’sk ( 44.029ºN 145.859ºE), 16. VI.1968, V. Richter, ( 1 ♂, ZISP); same place, 5.VII.1979, A. Zinov’ev ( 1 ♂, ZISP); Paramushir I., Severo-Kuril’sk ( 50.679ºN 156.132ºE), 16.VIII.1978, A. Zinov’ev ( 1 ♂, ZISP); Shikotan I., Malokuril’sk ( 43.869ºN 146.829ºE), 24.IX.1968, Gorodkov ( 28 ♂♂, 8 ♀♀, ZISP); Smolensk Oblast: “Smolenskoe poozer’e” National Park ( 55.511ºN 31.839ºE), 29, 30.VII. and 9.VIII.1991, Zlobin ( 2 ♂♂, 1 ♀, ZISP); Tyumen’ Oblast: “Elizarovskoe, Tobol. gub.” (=Elizarovo: 61.395ºN 68.295ºE), 7. VI.1907, M. Lagin ( 1 ♂, ZMUM); 75 km WSW of Samburg (ca. 67.035ºN 76.541ºE), 6–9.VIII.1976, Gorodkov ( 9 ♂♂, 8 ♀♀, ZISP); Labytnangi ( 66.657ºN 66.391ºE), 30. VI., 8–21.VII.1973, 28.VII.1981, V. Sychevskaya, G.A. Veselkin ( 11 ♂♂, 4 ♀♀, ZMUM); Cape Kamennyy ( 68.495ºN 73.582ºE), 9.VIII.1967, Gorodkov ( 3 ♂♂, 3 ♀♀, ZISP); Neroyka (ca. 64.57ºN 59.67ºE), 650 m, 6.IX.1990, Malozemov ( 1 ♂, ZISP); same place, 450 m, 29. VI.1990, Malozemov ( 1 ♂, ZISP); Salekhard ( 66.53ºN 66.613ºE), 3.VIII.1961, Gorodkov ( 4 ♂♂, 2 ♀♀, ZISP); same place, 4–5.VIII.1976, Gorodkov ( 3 ♂♂, 2 ♀♀, ZISP); Saranpaul’ ( 64.257ºN 60.917ºE), 23. VI.1989, Malozemov ( 2 ♂♂, ZISP); Tarko-Sale ( 64.914ºN 77.766ºE), 1 and 2.IX.1982, Gorodkov ( 4 ♂♂, 5 ♀♀, ZISP); Vologda Oblast: Vologda env. ( 59.207ºN 39.904ºE), 8.IX.1981, Gorodkov ( 1 ♀, ZISP); Yakutia: Aykhal ( 65.944ºN 111.495ºE), 19.VIII.1988, Gorodkov ( 1 ♂, 1 ♀, ZISP); Batagay ( 67.653ºN 134.635ºE), 31.VII.2008, A. Ovchinnikov ( 1 ♀, ZISP); Bestyakh ( 65.281ºN 124.120ºE), 14. VI.1912, Naumov ( 1 ♀, ZISP); Khaptagay ( 61.787ºN 129.795ºE), 30. VI.1974, Nartshuk ( 1 ♂, ZISP); left bank of the River Yana opposite of Verkhoyansk ( 67.55ºN 133.359ºE), 21 and 23.VII.1974, Nartshuk ( 5 ♂♂, 3 ♀♀, ZISP); Lensk ( 60.727ºN 114.943ºE), 12 and 14.IX.1987, Gorodkov ( 1 ♂, 1 ♀, ZISP); Mikhaylovka, 60 km NE of Amga ( 61.214ºN 132.681ºE), 15.VIII.1984, Averinskiy ( 5 ♂♂, ZISP); Mirnyy ( 62.540ºN 113.962ºE), 22.VIII.1988, Gorodkov ( 1 ♂, ZISP); Neryuktyayinsk-1 ( 60.258ºN 119.673ºE), 11.VII. 2008, A. Ovchinnikov ( 2 ♂♂, 1 ♀, ZISP); Nyurba ( 63.285ºN 118.331ºE), 18. VI.1987, Zlobin ( 3 ♂♂, 1 ♀, ZISP); Oktemtsy env. ( 61.672ºN 129.417ºE), 18.VII.1977, Bagachanova ( 1 ♀, ZISP); Olekminsk ( 60.370ºN 120.436ºE), 29.VIII.1988, Gorodkov ( 2 ♂♂, 1 ♀, ZISP); Olenek ( 68.504ºN 112.457ºE), 9.VIII.1973, Gorodkov ( 10 ♂♂, 2 ♀♀, ZISP); Oleneksky Bay, Ystannakh-Khocho ( 72.58ºN 121.42ºE), 20.VIII.2010, A. Yadrenkin ( 1 ♂, ZMUM); Pokrovsk ( 61.467ºN 129.101ºE), 21.VII.1986, Makarkin ( 2 ♂♂, IBSS); River Yana, Stolby env. ( 67.531ºN 134.087ºE), 29.VII. 2008, A. Ovchinnikov ( 6 ♂♂, 1 ♀, ZISP); Saskylakh env. ( 71.966ºN 114.094ºE), 9.VIII.1988, Gorodkov ( 11 ♂♂, 1 ♀, ZISP); Srednekolymsk ( 67.460ºN 153.710ºE), 16.VIII.1974, Gorodkov ( 6 ♂♂, 3 ♀♀, ZISP); Tiksi env. ( 71.635ºN 128.857ºE), 30.VII.1990, Kaspyaryan ( 2 ♀♀, ZISP); Tolon ( 59.458ºN 111.539ºE), 14.VII.1987, Zlobin ( 1 ♂, ZISP); Tommot ( 58.963ºN 126.282ºE), 23.VIII.1969, Sychevskaya ( 1 ♀, ZISP); Udachnyy ( 66.405ºN 112.2994ºE), 21.VIII.1988, Gorodkov ( 1 ♂, ZISP); Verkhoyansk ( 67.548ºN 133.396ºE), 3.VIII.1969, Sychevskaya ( 3 ♂♂, 1 ♀, ZISP); Vilyuysk env. ( 63.750ºN 121.637ºE), 24.VIII.1988, Gorodkov ( 1 ♀, ZISP); Yakutsk env. ( 62.034ºN 129.618ºE), 3.VIII.2008, A. Ovchinnikov ( 1 ♂, ZISP); Zhigansk ( 66.766ºN 123.372ºE), 7– 15.VIII.1973, Gorodkov ( 7 ♂♂, 5 ♀♀, ZISP); Zabaikalsky Krai: Chita ( 52.065ºN 113.444ºE), 10.IX.1974, Gorodkov ( 1 ♀, ZISP); Ingoda ( 51.851ºN 113.097ºE), 7.VII.1971, V. Richter ( 2 ♂♂, ZISP); Kyra ( 49.581ºN 111.974ºE), 17. VI.1975, V. Richter ( 1 ♂, ZISP); Olovyannaya ( 50.994ºN 115.561ºE), 9.VII.1971, V. Richter ( 1 ♂, ZISP). See also Ozerov & Krivosheina [2014: 218, as S. suilla ] and Ozerov & Barkalov [2014: 564, as S. suilla ].

DESCRIPTION. Male. Female. Body-length 3.8– 7.7 mm.

Head. Frontal vitta yellow or reddish-yellow, matt; fronto-orbital plate blackish, greyish dusted. Face, parafacial and gena yellow, with delicate whitish reflection. Postcranium black in upper third or half and yellow in lower part. 3 orbital and 3–4 frontal setae present. Antenna reddish-yellow. Postpedicel approximately twice as long as wide. Arista pubescent or plumose, the longest hairs approximately equal or more than 1/2 width of postpedicel. Palpus yellow.

Thorax greyish dusted. Scutum black completely, only postpronotal lobe usually yellowish; pleural sclerites yellowish in ground color, but anatergite, katatergite and mediotergite always black. Acrostichals setulose in two rows, but prescutellar pair stronger than other acrostichals, dorsocentrals 2+3, intra-alars 1+2, supra-alars 1+2, postpronotals 2. Katepisternum posteriorly with yellow hairs and setulae and, as a rule, without black setulae. Anepimeron bare. Scutellum black, greyish dusted, with a pair of strong basal scutellar and a pair of strong apical scutellar setae.

Legs yellow. Fore femur covered with hairs, but without conspicuous setae in male, with a row of thin dorsal setae in female. Fore tibia with 2 dorsal, 0–1 posterior, 1 preapical dorsal, and 1 posterior apical setae. Mid femur with 1 preapical seta in male and usually with a row of anterodorsal setae in female, also with 1 preapical posterior and 1 preapical posterodorsal setae in both sexes. Mid tibia with 1–2 posterodorsal, 1 anterodorsal setae and a ring of apicals. Hind femur without conspicuous setae in male and with a row of anterodorsal setae in female. Hind tibia with 2– 3 anterodorsal, 2–3 posterodorsal, 1 preapical dorsal, 1 preapical anterodorsal, 1 apical anteroventral setae.

Wing tinged with brownish; crossveins r-m and dm-cu darkened.

Abdomen from yellowish to dark brownish, delicately greyish dusted, covered with not long hairs; female tergites with black marginal setulae. Female tergite 7 shiny. Male sternite 4 approximately 2 times as long as wide (Fig. 37); sternite 5 with a pair of narrow median processes ( Fig. 112 View Figs 106–114 ). Epandrium, cercal plate and surstyli as in Figs 113, 114 View Figs 106–114 .

DISTRIBUTION. Russia: throughout midland, north to southern tundra. — Europe (widespread), China, Japan, Kazakhstan, Mongolia, Syria [ Šifner, 2008: 172], North America; a widespread and common species, but more abundant in the forest zone.