Chimalfus, Larsen & Solodovnikov & Reyes-Hernández, 2025

3.1.1. Chimalfus gen. nov

Fig. 1A–C View Fig , 2A -C View Fig , 3 View Fig , 4 View Fig , 5 View Fig and 6 View Fig .

Type species: Heterothops discoideus Fairmaire and Germain, 1862 , here designated.

Included species: Chimalfus brevis ( S´aiz, 1971) comb. nov., Chimalfus brunneus sp. nov., and Chimalfus discoideus ( Fairmaire and Germain, 1862) comb. nov.

Diagnosis: Chimalfus can be distinguished from all other genera in Amblyopinina based on the combination of the following characters: anterior postgenal ridge present (APGR; Fig. 5C View Fig ); basisternum without pair of macrosetae; pronotum with rounded flexible translucent postcoxal hypomeral process; middle and hind legs without black combs or pale adhesive setae; two long or medium size empodial setae on all tarsi; basal portion of metacoxa without proximal transverse carina; abdomen with protergal glands; all tergites without posterior transverse basal carina, pattern of posteromarginal macrosetae (not counting the paratergites) on each side of tergites III to VI: 1-2-3-3.

Description: Head about as wide as long (HW/HL 0.9–1.1); with complete frontoclypeal (epistomal) suture (junction of the anterior tentorial arms visible through the cuticle externally); disc without either coarse non-setiferous punctures or distinct concavity between antennal insertions; neck moderately wide (NW/HW 0.75–0.90); dorsal macrosetae ( Fig. 1 View Fig ) represented by frontoclypeal (landmarks 1 and 2), anterolateral clypeal, anterior frontal (landmarks 3 and 4), parocular (single; landmarks 5 and 6), posterior frontal (landmarks 7 and 8), temporal (landmarks 9 and 10), and single basal (landmarks 11 and 12) punctures; supra-antennal, interocular, and dorsogenal punctures absent; basal punctures located distinctly posterior to posterior frontal punctures, posterior frontal punctures located anterior to temporal punctures; on ventral side postocular punctures present, but infraorbital absent; submentum with one pair of macrosetae; mentum with seta alpha present and seta beta absent; ventral basal, postgenal, postmandibular ridges present; ventral basal ridge joining gular sutures; postmandibular ridge (in ventral view) extended parallel and close to eye margin; infraorbital ridge merged with postgenal ridge; nuchal ridge present dorsally and laterally; EYL/HL ratio between ½ and ¾; gular sutures moderately separated from each other; anterior postgenal ridge present ( Fig. 5C View Fig ); gula with distinct transverse basal impression. Antennae inserted at equal distance or closer to eye than to apical margin of frontoclypeus; tomentose pubescence begins on fourth antennomere (a4), and as dense as in the following antennomeres; a11 with subapical rounded field and lateral pits; a3 not visibly longer than a2 (a3/a2 < 1.1); a1 distinctly shorter than a3 and a2 combined; a11 moderately longer than a10 (a11/a10 ≤ 1.5). Mandibles with dorso-lateral groove; in dorsal view with straight external edge, with both base and apex gently curved; in lateral view apical portion of mandible deflexed ventrally at an angle of 25–30 ◦; on right mandible space between proximal and distal teeth (not bifid) narrow and smooth, left mandible with non bifid proximal tooth only. Labrum with wide transparent apical membrane, anterior margin of heavier sclerotized part entire, without emargination. Maxillary and labial palps with apical segment having evenly narrowed apex so its shape close to subconical; maxillary palpomere 4 (mp4, apical) distinctly shorter than mp2 (mp4/ mp2 length ratio ≤ 0.75); labial palpomere 2 (lp2, preapical), markedly more dilated compared to lp3 (apical) (lp3/lp2 width < 0.75).

Pronotum slightly transverse (PW/PL > 1.1); with one pair of punctures in dorsal series, located far from anterior margin of pronotum; sublateral setiferous punctures absent; basisternum triangular with lateral arms narrowed at least subapically and disc (in lateral view) protruding medially, without pair of macrosetae; antesternal membrane without distinct semi-sclerotized patch; sternacostal ridge (transverse carina) medially protruding (better seen from lateral view); probasisternum distinctly longer than profurcasternum; profurcasternum with apophysis; pronotum and prosternum not fused in procoxal cavity, pronotosternal suture complete; pronotal hypomeron with flexible postcoxal process.

Legs with 5-5-5 tarsal formula, apical tarsomere without dorsal setae but with pair of long empodial setae; protarsomeres 1–3 broad with pale setae; profemur with apical row of lateroventral spines reaching halfway its apex; mesotarsi without pale adhesive setae or black combs; metafemur with row of laterodorsal setae; metacoxae with spines on its ventral posterolateral lobe; metakatepisternal processes divided; metacoxae with basal part distinctly wider and more convex than apical part.

Elytra without humeral spines or spine-like setae, with subbasal ridge, with setiferous punctation on disc and epipleuron even; mesoscutellum without posterior scutellar ridge; mesanapleural sutures distinctly oblique and fused with prepectus; apex of intercoxal process narrowly pointed, forming sharp (acute) angle; mesofurcasternal apophysis projected towards mesosternacostal suture medially; mesocoxal cavities contiguous; pericoxal ridge complete. Metaventral submarginal ridge long (parallel along most of metaventral dorsal edge); basal portion of each metacoxa without a transverse ridge.

Abdomen tapered apicad; first tergite (reduced, hidden under elytra) with protergal glands rounded to elliptical, less than three times as long as wide; tergites III-VII without posterior transverse basal carina; tergite VII with anterior transverse basal carina not continuing to paratergites; pattern of posteromarginal macrosetae on each side of tergites III to VI 1- 2-3-3: tergite III with one macroseta on each side, tergite IV with two, and tergites V and VI each with three macrosetae on each side; tergites III-VI with apical row of setae at distinctly crenulate apical margin; tergites III-IV with punctation on disc consisting of fine to moderate impressions and with setiferous punctation forming patches of pale radiating setae; tergite VIII longer than tergite IX; tergite VIII without medial apical emargination; tergite X in males emarginate medio-apically; female tergite X emarginate medio-apically; sternite III with basal transverse carina laterally converging abruptly at an acute angle with rounded tip; male sternite IX slender pear-shaped with asymmetrical basal portion, without distinct median emargination apically; lateral tergal sclerites IX inflated. Females with second gonocoxite having one macroseta.

Aedeagus: paramere without sensory peg setae, flat, at base fused to median lobe and very closely appressed to it along its entire length; apex of paramere strongly produced over apex of median lobe.

Distribution. The genus is confined to central and southern Chile ( Fig. 6 View Fig ).

Etymology. The name is a latinization of the Mapudungun word “Chimalfe,” which in mapuche language means “dragonfly” and refers to the large eyes in this genus. Gender masculine.

Bionomics. The species of Chimalfus were collected at elevations ranging from 10 to 1700 m above sea level. Species were primarily collected from forest leaf and log litter, moss and other debris through sifting. Additional specimens were obtained using pitfall traps (baited with various substances such as carrion, dung and banana), and flight intercept traps. They were usually confined to very moist forest leaf litter, boggy mixed forest remnants, and Valdivian rainforest with thick bamboo understory.